Primulaceae

Primulaceae

Primula gracilipes

Primulaceae

Androsace rotundifolia

Primulaceae

Primula capitata

Primulaceae

Primula irregularis

Primulaceae

Primulaceae

Primula obliqua

by GP Jürgens

W. W. Sm.

BhutanPG2011 409
Bhutan Jule-Tso, 4300m June

Description

Primula obliqua W. W. Sm. is a robust alpine primrose of the eastern Himalaya, recorded by GBIF from Nepal, India, Bhutan and China across some 183 occurrences. It is a plant of the upper alpine zone, growing on cool, humid open slopes, rocky meadows, moraine and the margins of melting snow patches, generally between 3800 and 4800 m. The plant photographed at Jule-Tso in Bhutan at about 4300 m in June, when the snow has just retreated, illustrates its preference for high, well-watered sites where it flowers early in the short alpine summer.

The species forms a stout perennial rosette of large, oblong to oblanceolate leaves that are somewhat fleshy and conspicuously dusted, especially when young, with pale yellowish or whitish farina on the undersurface and stem. The leaves taper to a broadly winged petiole and have finely toothed margins. From the centre rises a stout scape bearing an umbel of several nodding flowers; the corolla is broadly campanulate, pale yellow to cream or greenish-yellow, with a short tube and spreading lobes, and the calyx and pedicels are mealy. The flowers are heterostylous, and the fruit is a capsule shorter than to about equalling the calyx.

Primula obliqua is classified in section Crystallophlomis (Nivales), a group of large-flowered, often farinose high-alpine primulas centred on the Sino-Himalayan mountains. It is distinguished within the section by its drooping, pale yellowish, mealy flowers. With a restricted but apparently stable range in remote high terrain, it is not currently regarded as threatened, though as a snow-bed and high-alpine specialist it is potentially vulnerable to climate-driven changes in snow cover. It exemplifies the rich endemic radiation of Primula along the eastern Himalayan arc.

Author: W. W. Sm. · Sources: POWO (Kew), GBIF, Flora of Bhutan, Flora of China, Richards monograph Primula

Primulaceae

Primula petiolaris

by GP Jürgens

Wall.

Nepal 95 108 P193 ergebnis
Nepal below Rupien La 3050m October
Garh 07 103 ergebnis
India Garhwal Meggu Kichatti 3100m, June
Garh 07 102 ergebnis
India Garhwal Meggu Kichatti 3100m, June

Description

Primula petiolaris Wall. is a small, cushion- or mat-forming primrose of the Himalaya, distributed through Nepal, northern India, Bhutan and south-west China. It is a plant of cool, moist, shaded sites in the upper-temperate to alpine zone, growing on humus-rich banks, in rock crevices, beside streams and on mossy ledges, typically around 3000 to 4000 m. The recorded gatherings from below the Rupien La in Nepal at 3050 m and from Meggu Kichatti in Garhwal at 3100 m, the latter flowering in June soon after snowmelt, illustrate its early-season flowering in damp, semi-shaded high-mountain habitats where it forms low rosettes among rocks and moss.

The plant is a low, tufted perennial forming dense rosettes of obovate to spathulate leaves that taper gradually into a winged petiole (the basis of the epithet petiolaris), with finely toothed margins and a somewhat fleshy texture. The flowers are borne singly or in small clusters nestled within or just above the foliage on very short scapes, so that the plant appears almost stemless at flowering. Each flower has a tubular calyx and a salver-shaped corolla, rose-pink to purple with a contrasting white or yellowish eye ringing the throat. As in many primulas the species is heterostylous, with pin and thrum flower forms, and sets a small capsule releasing numerous fine seeds.

Primula petiolaris belongs to the large Himalayan radiation of section Petiolares, an intricate group of early-flowering, rosette-forming primroses whose species delimitation has long challenged taxonomists. Its precocious flowering, often as snow recedes, points to early-emerging insects as pollinators. Choice plants of this section are prized by alpine-garden growers, but P. petiolaris remains widespread and is not regarded as threatened, though its specialised montane microhabitats are sensitive to disturbance and climatic change.

Author: Wall. · Sources: POWO (Kew), GBIF, Flora of China (Primulaceae), Annotated Checklist of the Flowering Plants of Nepal, Flora of Bhutan, The Genus Primula (Richards)

Primulaceae

Primula prenantha

by G & P Jürgens

Balf.f. & W.W.Sm.

Sikkim 2006 435 ergebnis
Sikkim 2006 361 ergebnis
India West-Bengal, Tamrong 2750m, May

Description

Primula prenantha is a small perennial herb distributed across the eastern Himalaya and adjacent mountains, with GBIF recording it from China, India, Myanmar, Nepal and Bhutan across some 93 georeferenced occurrences. It is a plant of moist, cool montane habitats, favouring stream margins, seepage slopes, wet meadows and the edges of evergreen and mixed broad-leaved forest, typically between about 2400 and 3600 m. The photographed population at Tamrong in West Bengal, at roughly 2750 m, flowering in May, is characteristic of this damp temperate belt where snowmelt keeps the ground saturated through the early growing season.

The plant forms a small basal rosette arising from a short rhizome with fibrous roots. The leaves are obovate to oblong-spathulate, narrowing gradually into a winged petiole, with a crenate to denticulate margin and an obtuse apex; the lamina is thin, glabrous and not farinose. From the rosette rises a slender erect scape bearing a terminal umbel of few to several nodding flowers subtended by small linear bracts. The flowers are heterostylous; the calyx is tubular-campanulate and lobed, and the corolla is yellow with a narrow tube and spreading, often emarginate lobes forming a flat or slightly cupped limb. The fruit is a small cylindrical capsule that dehisces by apical valves to release numerous minute seeds.

The species belongs to Primula section Sikkimensis (in the broad sense, allied to the yellow-flowered Petiolares-Sikkimensis assemblage), a group centred on the Sino-Himalayan region whose diversification tracks the uplift of the Himalaya and the associated humid monsoon gradients. Its dependence on cool, perennially moist microhabitats makes it sensitive to changes in snowpack and montane hydrology, and like many Sino-Himalayan primulas it is locally rather than broadly distributed. It is not currently regarded as globally threatened, but its narrow ecological tolerance warrants attention under warming and drying climatic scenarios.

Author: Balf.f. & W.W.Sm. · Sources: POWO (Kew), GBIF, Flora of China, Flora of Bhutan, Annotated Checklist of the Flowering Plants of Nepal, John Richards "Primula" (monograph)

Primulaceae

Primula primulina

by GP Jürgens

(Spreng.) Hara

BhutanPG2011 625 01
Bhutan Ludlow-Sherriff-Trail Shashuna-Dem-Tso, 3900m June
Sikkim2003 327 P464
Sikkim Saniti-Lake 4200m August

Description

Primula primulina is a dwarf alpine primrose of the Himalayan high country, distributed through Nepal, northern India, Bhutan and southern China, with several hundred georeferenced occurrences recorded in GBIF. It is a cushion- or mat-forming species of the upper alpine and nival zones, growing on open stony slopes, moist alpine turf, rock ledges and the margins of high-elevation tarns kept moist by late snowmelt. The material from the Ludlow-Sherriff Trail at Shashuna-Dem-Tso in Bhutan (about 3900 m, June) and from Saniti Lake in Sikkim (4200 m, August) reflects its association with glacial lake basins and its summer flowering across the monsoon months.

P. primulina is a diminutive, densely tufted perennial forming low rosettes or compact cushions from a slender branching rootstock. The leaves are small, oblanceolate to spathulate, often densely crowded, with an entire or finely toothed margin and frequently bearing farina, the rosettes sometimes marcescent at the base. The flowers are borne singly or few together on very short scapes scarcely exceeding the foliage, giving an almost sessile appearance. The corolla is salverform with a slender tube and spreading, notched lobes, typically rose-purple to pink with a paler or yellowish eye; the calyx is tubular and lobed. The fruit is a small capsule enclosed by or barely exceeding the persistent calyx.

The species is placed in Primula section Minutissimae, the group of minute, cushion-forming Sino-Himalayan primroses to which P. primulina lends its name as a characteristic member. Long-established synonymy reflects its complex nomenclatural history under several earlier basionyms. As a specialist of high-altitude snowbed and lakeshore microhabitats, it is closely tied to persistent snow cover and is therefore among the alpine taxa most likely to be affected by warming-driven contraction of suitable habitat, though it currently carries no formal conservation assessment.

Author: (Spreng.) Hara · Sources: POWO (Kew), GBIF, Flora of Bhutan, Flora of China, The Genus Primula (Richards)

Primulaceae

Primula pulchra

by GP Jürgens

G.Watt

Sikkim2004 142 G244
Sikkim/India Singalila 3800m April

Description

Primula pulchra is an eastern Himalayan species distributed across Nepal, India, Bhutan, southern China (Xizang, Yunnan) and northern Myanmar, with GBIF documenting some 39 georeferenced occurrences across this comparatively narrow range. It is a plant of the upper temperate and subalpine zones, typically growing between about 3000 and 4300 m on moist, humus-rich slopes, in open Rhododendron scrub, along stream margins and on grazed alpine meadows that remain snow-covered for much of the winter. The photographed material from the Singalila Ridge on the Sikkim–West Bengal border at roughly 3800 m, flowering in April, is characteristic: the species is among the earliest primulas to bloom as the snowmelt recedes.

It is a small, rosette-forming perennial arising from a short rhizome with fibrous roots. The leaves are obovate to oblong-spathulate, narrowed to a winged petiole, the margin crenate-dentate, and the lower surface typically clad in a pale to yellowish farina. The scape carries a one- to few-flowered umbel subtended by short bracts. Flowers are heterostylous and showy, the corolla rose-purple to violet with a pale or yellow eye, the lobes obcordate and emarginate; the calyx is campanulate and often farinose. The fruit is a small capsule about equalling the persistent calyx, releasing numerous angular seeds.

Primula pulchra belongs to section Petiolares, a large, taxonomically intricate group of Sino-Himalayan primulas characterised by early flowering and resting-bud overwintering. Within it the species is allied to P. calderiana and P. boothii, from which it is separated by leaf shape, farina distribution and corolla detail; intergradation has complicated its delimitation. As a snow-bed and meadow specialist it is sensitive to grazing and to shifts in snow-lie driven by climate warming, though its presence in several protected high-altitude areas affords some security.

Author: G.Watt · Sources: POWO (Kew), GBIF, Flora of China, Flora of Bhutan, Smith & Fletcher monograph of the genus Primula, Richards 'Primula' (2003)

Primulaceae

Primula reidii

by G & P Jürgens

Duthie

Nepal 99 515 ergebnis
West-Nepal Churta-Khola 3800m, June
Nepal 99 804 ergebnis
West-Nepal Baga-La-Phedi 4350m, July
Nepal 99 461 ergebnis
West-Nepal Naphukana 2830m, June

Description

Primula reidii is a celebrated alpine primrose of the western and central Himalaya, with about 80 GBIF occurrences recorded from Nepal, India and Pakistan. It is a plant of cool, humid high-elevation habitats, growing among moist rocks, on shaded ledges, in crevices and on steep banks where seepage and snowmelt maintain constant moisture. The documented localities span a wide altitudinal range, from Naphukana in West Nepal at about 2830 m and Churta-Khola at 3800 m in June to Baga-La-Phedi at around 4350 m in July, confirming its occurrence from the upper temperate well into the alpine zone during the early monsoon flowering season.

It is a perennial rosette herb, the leaves clustered at the base into a rosette, oblong to oblanceolate, soft-textured, crenate to toothed, and frequently bearing a white or yellowish farinose (mealy) coating on the scape, calyx and leaf undersides. The flowers are borne in a terminal umbel or short head on a leafless scape, subtended by an involucre; each has a tubular, farinose, five-lobed calyx and a bell-shaped (campanulate), nodding corolla. The famously fragrant flowers are white to pale blue with five rounded, often notched lobes. The species is typically homostylous, and the fruit is a small capsule releasing numerous fine seeds.

The genus Primula reaches its greatest diversity in the Sino-Himalayan mountains, and P. reidii, described by Duthie, belongs to the bell-flowered alpine assemblage traditionally placed in section Soldanelloides, esteemed by horticulturists for its scented, nodding bells. It is a much-admired but somewhat localised alpine restricted to specific moist rock habitats; while not formally assessed as threatened, such niche-restricted alpines are sensitive to collection pressure, grazing and the warming-driven desiccation of high-altitude seeps.

Author: Duthie · Sources: POWO (Kew), GBIF, Flora of Nepal, Annotated Checklist of the Flowering Plants of Nepal, Richards Primula, Smith & Fletcher revision of Primula

Primulaceae

Primula rosea

by G & P Jürgens

Royle

Garh 07 407 ergebnis
India Garhwal Khiar Kutti 4000m, June

Description

Primula rosea is a western Himalayan alpine endemic distributed through northern India, Pakistan, Afghanistan and adjacent China, with GBIF recording around 255 occurrences, supplemented by numerous cultivated records in Europe and North America where it is grown as an ornamental. It is a true marsh-dweller of the high alpine zone, generally between roughly 3000 and 4300 m, the photographed plant coming from Khiar Kutti in Indian Garhwal at 4000 m, flowering in June at snowmelt. It grows in boggy alpine meadows, beside streams and springs, in seepage flushes and on wet turf, often forming spectacular rose-pink colonies as the snow recedes.

The species is a small, tufted, rosette-forming perennial herb with short rhizomes, the basal leaves developing fully only after flowering. These are oblong-obovate to oblanceolate, narrowed to a winged petiole, finely toothed and bright green, more or less glabrous and lacking the mealy farina of some congeners. The leafless scape bears an umbel of a few to several flowers; each flower is rich rose-pink to carmine with a contrasting yellow eye, the corolla salverform with a slender tube and spreading, notched lobes, and the species is heterostylous (pin and thrum). The fruit is a small capsule shedding numerous seeds.

Primula rosea belongs to section Aleuritia (the farinose primroses) within the large Holarctic genus Primula, though it is itself essentially efarinose. Its heterostyly enforces outcrossing by insect pollinators active in the brief alpine season. Locally abundant in suitable wet habitats and not globally threatened, it is nonetheless a habitat specialist whose marshy alpine flushes are sensitive to grazing, trampling and hydrological change.

Author: Royle · Sources: POWO (Kew), GBIF, Flora of Pakistan, Flora Simlensis, Flora of British India (Hooker), Richards Primula (monograph), Smith & Fletcher The Genus Primula

Primulaceae

Primula sapphirina

by GP Jürgens

Hook.f. & Thomson ex. G.Watt

BhutanPG2011 299
Bhutan Jule-Tso, 4000m June
Nepal 99 688 1 ergebnis
West-Nepal, Pungmo-Khola, July

Description

Primula sapphirina is a diminutive alpine primrose of the high Eastern Himalaya, recorded from India, Bhutan, Nepal and south-western China, with GBIF documenting around 80 occurrences from this restricted high-altitude range. It is confined to the alpine zone, growing on moist turf, mossy banks, seepage slopes and the edges of melting snow patches where the ground stays cool and damp through the brief growing season. The cited localities underline its elevation: Jule-Tso in Bhutan at about 4000 m flowering in June and the Pungmo-Khola in West Nepal in July, both well above treeline, where it flowers as the snow recedes and the short subnival summer begins.

It is a small, delicate perennial herb forming a slender, few-leaved tuft from a short rootstock, often farinose. The leaves are basal, small, spathulate to oblong, narrowed into a winged petiole, with finely toothed or crenate margins. The slender scape bears few small flowers, sometimes solitary, nodding on capillary pedicels; the calyx is tubular-campanulate and the corolla is a clear sapphire-blue to violet, the limb shallowly lobed with a narrow tube, giving the plant its epithet. The species is typically heterostylous with pin and thrum forms. The fruit is a small membranous capsule containing numerous minute seeds.

Primula sapphirina, attributed to Hooker f. & Thomson ex G. Watt, belongs to section Minutissimae (the so-called "bird's-eye" alpine primulas), a group of tiny high-elevation species characterised by farinose, cushion-like habit and small blue flowers. As an obligate alpine, it depends on snowmelt moisture and stable turf, and its pollination by small flies and bees occurs within a narrow window. Its restricted distribution and specialised habitat make it sensitive to climate-driven upslope shifts, though no formal conservation assessment has yet been published.

Author: Hook.f. & Thomson ex. G.Watt · Sources: POWO (Kew), GBIF, Flora of Bhutan, Flora of China, Annotated Checklist of the Flowering Plants of Nepal, Richards' monograph Primula

Primulaceae

Primula scapigera

by GP Jürgens

Craib

Sikkim2004 75 P131
Sikkim West-Bengal, Bersay 2600m April
Sikkim2004 313 G563
Sikkim/India Kokchurong 3600m May
Sikkim2004 314 G564
Sikkim/India Kokchurong 3600m May

Description

Primula scapigera is an eastern Himalayan primrose recorded from northern India (Sikkim, West Bengal), Nepal, Bhutan and adjacent China, with 67 GBIF occurrences also reflecting cultivated material reaching Japan. It grows in cool, humid montane and subalpine habitats, on mossy banks, in open Rhododendron and conifer forest and along streamsides, generally between 2500 and 3800 m. The illustrated plants were collected at Bersay in the Sikkim/West Bengal borderland at 2600 m in April and at Kokchurong in Sikkim at 3600 m in May, spanning the species' spring to early-summer flowering across its altitudinal range.

It is a small perennial herb forming neat rosettes of obovate to spathulate leaves that are crenate to denticulate at the margin, narrowed to a winged petiole and often coated beneath, like the scape and calyx, with pale farina. The slender erect scape bears a terminal umbel of a few to several flowers. The corolla is rose-pink to purplish with a contrasting yellow or whitish eye, the limb divided into five obcordate, notched lobes; the plants are heterostylous, producing pin and thrum morphs. The fruit is a small capsule shorter than or equalling the persistent calyx.

The species is placed in Primula section Petiolares, a largely Sino-Himalayan group of farinose, rosette-forming primroses centred on the eastern Himalaya and Hengduan ranges. Within the section it is allied to P. petiolaris and P. gracilipes, from which it is separated by its scapose inflorescence and leaf characters. Like its congeners it depends on insect pollination facilitated by its distyly, and its restricted high-rainfall habitat makes it sensitive to disturbance and warming. Though not formally assessed, it remains a localised eastern Himalayan endemic of conservation interest.

Author: Craib · Sources: POWO (Kew), GBIF, Flora of Bhutan, Flora of China, Richards: Primula

Primulaceae

Primula sharmae

by G & P Jürgens

H.R.Fletcher

Nepal 99 575 ergebnis
West-Nepal Bheri-Khola 2850m, July
Nepal 99 574 ergebnis
West-Nepal Bheri-Khola 2850m, July

Description

Primula sharmae is a narrowly distributed primrose apparently endemic to Nepal, where it is the sole region documented in GBIF among its roughly 72 occurrence records. It is a plant of cool, humid montane environments, occupying moist rocky slopes, shaded banks, ledges and the margins of streams and forest in the temperate Himalayan zone. The population photographed along the Bheri-Khola in West Nepal at about 2850 m in July was recorded during the monsoon, the principal flowering and growth period when persistent cloud and rainfall keep these slopes saturated and favour the moisture-demanding primulas.

It is a perennial rosette herb, the leaves crowded at the base into a basal rosette, oblong to oblanceolate or spathulate, with toothed or crenate margins and gradually tapering into a winged petiole; the foliage and scape may bear a fine farinose (mealy) covering characteristic of many Himalayan primulas. The flowers are borne in a terminal umbel atop a leafless scape, subtended by an involucre of bracts; each flower has a tubular, five-lobed calyx and a salverform corolla with a slender tube and five spreading, often notched (emarginate) lobes. The species is heterostylous, presenting pin and thrum floral morphs, and the fruit is a small dehiscent capsule releasing numerous seeds.

The genus Primula is exceptionally diverse in the Himalaya, where many species are narrow endemics confined to particular valley systems, and P. sharmae, described by H.R.Fletcher, is one such restricted-range taxon. Endemic primulas with limited known distributions are of inherent conservation interest, as small populations tied to specific moist microhabitats are vulnerable to disturbance, grazing and climatic drying. Continued field documentation across western Nepal would help clarify its true range and population status.

Author: H.R.Fletcher · Sources: POWO (Kew), GBIF, Flora of Nepal, Annotated Checklist of the Flowering Plants of Nepal, Richards Primula, Smith & Fletcher revision of Primula