Asteraceae

Asteraceae

Ajania myriantha

Asteraceae

Anaphalis busua

Asteraceae

Aster peduncularis 

Asteraceae

Aster ageratoides

Asteraceae

Asteraceae

Anaphalis nepalensis

by GP Jürgens

(Spreng.) Hand.-Mazz.

Garh 07 207 ergebnis
India Garhwal Panwali 3300m, June

Description

Anaphalis nepalensis is a widespread Sino-Himalayan everlasting distributed across China, Taiwan, Nepal, India, Pakistan, Bhutan, Afghanistan and Myanmar, with over 3600 georeferenced records making it one of the better-documented Himalayan composites. It occupies open alpine and subalpine meadows, grassy slopes, rocky pastures and forest clearings, generally between about 3000 and 4500 m. The photographed plant from Panwali in the Garhwal Himalaya at 3300 m, flowering in June, typifies its grassy mid-alpine habitat where it forms low cushions or spreading patches in short turf.

It is a perennial, white-woolly herb arising from a creeping rhizome, with prostrate to ascending stems clothed in dense grey-white tomentum. The leaves are oblanceolate to spathulate, often crowded in a basal rosette and reducing upward, cottony on both surfaces. The capitula are arranged singly or in small terminal clusters and are dioecious or subdioecious; the conspicuous part is the spreading ring of dry, papery, pearly-white involucral bracts surrounding the small yellowish disc florets, an arrangement that gives the genus its "pearly everlasting" character. The achenes bear a pappus of fine barbellate bristles.

The persistent, scarious bracts retain their form long after flowering, an adaptation suiting the plant to harsh, exposed alpine conditions and aiding identification. Anaphalis nepalensis is a variable species with several recognised varieties differing in stature and leaf form, and it is sometimes confused with related Himalayan congeners. Common and ecologically adaptable, it is not of conservation concern and is occasionally grown as a rock-garden ornamental.

Author: (Spreng.) Hand.-Mazz. · Sources: POWO (Kew), GBIF, Flora of China, Annotated Checklist of the Flowering Plants of Nepal, Compositae monographs (Handel-Mazzetti)

Asteraceae

Anaphalis nepalensis var. monocephala

by GP Jürgens

(DC.) Hand.-Mazz.

Sikkim2003 187 P255
Sikkim Dzongri 4000m 08.2003
Nepal 99 274 ergebnis
West-Nepal Loccu 3800m, June
Garw 2005 150 ergebnis ergebnis
India Garhwal Kuari-Pass 3900m, September

Description

Anaphalis nepalensis var. monocephala is a high-alpine pearly everlasting of the Himalaya and adjacent China, recorded from Nepal, India, Bhutan, Pakistan, Taiwan and south-west China across 426 georeferenced occurrences. It occupies open alpine turf, screes, rocky slopes and gravelly moraine in the upper alpine zone. The cited collections at Dzongri in Sikkim (4000 m), Loccu in west Nepal (3800 m) and the Kuari Pass in Garhwal (3900 m) confirm its preference for elevations of roughly 3500-4800 m, where it flowers through the short summer and into early autumn. The variety is distinguished from typical A. nepalensis chiefly by its solitary or few terminal capitula on dwarf, cushion-forming stems.

It is a low, mat- or cushion-forming perennial herb with a branched woody caudex and short, white-woolly (lanate) stems and foliage. The leaves are spathulate to oblanceolate, crowded toward the base, densely grey-tomentose and frequently bearing a tuft of wool at the apex. The capitula are usually solitary (monocephalous), comparatively large, and surrounded by numerous papery, pearly-white, radiating involucral bracts that simulate ray florets; the true florets are small and yellowish, the plant being functionally dioecious. The fruit is a tiny cypsela crowned by a pappus of barbellate bristles.

The genus Anaphalis belongs to tribe Gnaphalieae (the cudweeds and everlastings), in which dry, persistent scarious bracts give the inflorescences their lasting quality. The white bracts and clustered florets attract small flies and bees, while the plumed cypselae are wind-dispersed. The variety is locally frequent in high Himalayan pastures and is not considered threatened, though sensitive to heavy grazing and trampling.

Author: (DC.) Hand.-Mazz. · Sources: POWO (Kew), GBIF, Flora of Bhutan, Flora of China, Flora of Nepal, Hilliard & Burtt Gnaphalieae studies

Asteraceae

Anaphalis royleana

by G & P Jürgens

DC.

Garw 2005 297 ergebnis
India Garhwal Patanathi 3750m, October
Garw 2005 50 ergebnis

Description

Anaphalis royleana is a high-montane to subalpine perennial widely distributed across the Sino-Himalayan region, recorded from India, Nepal, Pakistan, Bhutan, Bangladesh, Myanmar, China and Taiwan. With 482 georeferenced occurrences, it is one of the more frequently collected members of the genus, occupying open stony slopes, alpine meadows, grazed pastures and the margins of rhododendron and birch scrub. The species typically ranges between roughly 3000 and 4500 m; the plant photographed at Patanathi in Garhwal, India, at 3750 m in October exemplifies its preference for well-drained, sunny situations where it flowers and fruits late into the season.

The plant forms tufted clumps from a woody rootstock, with erect, simple, white-woolly (lanate) stems. The leaves are alternate, sessile and frequently decurrent, linear to oblong-lanceolate, entire-margined, and densely grey- to white-tomentose, the lower surface usually more persistently floccose than the upper. The capitula are small, numerous, and aggregated into dense terminal corymbs. Each head is subtended by several rows of dry, papery (scarious), pearly-white involucral bracts that give the genus its garden name; the florets are functionally unisexual, the central disc florets staminate and the marginal florets pistillate. The fruit is a small cylindrical cypsela crowned by a pappus of slender, barbellate bristles facilitating wind dispersal.

Anaphalis is a large Old World genus closely allied to Helichrysum and Gnaphalium within the tribe Gnaphalieae, characterised by its dioecious or subdioecious capitula. A. royleana belongs to a taxonomically intricate complex of Himalayan everlastings in which leaf indumentum and bract form provide the principal diagnostic characters. Its broad elevational and geographic amplitude, tolerance of grazing, and effective anemochory render it secure, and it is not considered threatened.

Author: DC. · Sources: POWO (Kew), GBIF, Flora of China, Flora of Bhutan, Flowers of the Himalaya (Polunin & Stainton), Anaphalis revisionary studies in Gnaphalieae

Asteraceae

Anaphalis triplinervis 

by GP Jürgens

Sims ex C.B.Clarke

Bhutan2015 408
Bhutan Tumching-La 4000m, October
Garw 2005 296 ergebnis
India Garhwal Kunol 2900m, September

Description

Anaphalis triplinervis is a widespread Himalayan everlasting recorded natively from Nepal, India, China, Pakistan, Bhutan, Myanmar and Afghanistan, and widely grown in gardens (hence European and North American records); GBIF holds over 900 occurrences. It is a plant of open subalpine and alpine slopes, grassy meadows, forest clearings and stony banks, ranging from about 2000 to over 4000 m. The photographed material is from Tumching-La in Bhutan at 4000 m in October and from Kunol in Garhwal, India, at 2900 m in September, spanning the autumn flowering across the central and eastern Himalaya.

The species is a tufted perennial herb with creeping rhizomes and erect, white-woolly stems. The leaves are alternate, oblong to obovate-lanceolate, sessile and often somewhat decurrent, with the three prominent longitudinal nerves that give the epithet "triplinervis"; they are green and thinly cobwebby above and densely white-tomentose beneath. The capitula are small and crowded in dense terminal corymbs, each surrounded by spreading, papery, pearly-white involucral bracts that form the showy "flower"; the true florets are small and yellowish, the plants typically dioecious or functionally so. The fruit is a tiny cypsela with a pappus of barbellate bristles.

Anaphalis belongs to the tribe Gnaphalieae, the cudweeds, and A. triplinervis is among the most familiar Himalayan species, its persistent papery bracts making it a popular everlasting for drying. Abundant and ecologically tolerant of grazed and disturbed slopes, it is of no conservation concern.

Author: Sims ex C.B.Clarke · Sources: POWO (Kew), GBIF, Flora of Bhutan, Flora of China, Annotated Checklist of the Flowering Plants of Nepal, Revision of Asiatic Anaphalis (Gnaphalieae)

Asteraceae

Anaphalis xylorhiza

by G & P Jürgens

Sch.Bip. ex Hook.f.

Sikkim2003 260 P358
Sikkim Dzongri 4000m, July

Description

Anaphalis xylorhiza is a high-alpine pearly-everlasting of the eastern Himalaya, recorded from China, Nepal, India and Bhutan, with about 416 georeferenced occurrences indicating a fairly frequent species along the inner high ranges. It grows in alpine meadows, on stony slopes, screes and rocky ridges above the treeline, typically between roughly 3500 and 4800 m. The plant was photographed at Dzongri in Sikkim at 4000 m in July, a setting representative of its ecology, flowering at the height of the monsoon on open, well-drained, frequently grazed alpine ground.

As the epithet xylorhiza indicates, it is a perennial herb with a thick, woody rootstock from which arise low, white-woolly stems. The leaves are linear to narrowly oblong or spathulate, often crowded toward the base, grey-green and clothed in a dense persistent felt of appressed hairs on both surfaces, the bases sometimes decurrent. The capitula are small and aggregated into compact terminal corymbose clusters; each head is surrounded by dry, papery, pearly-white involucral bracts that give the genus its everlasting quality. The plants are essentially dioecious, with male and female heads on separate individuals, and the fruit is a minute cypsela bearing a pappus of slender bristles.

Anaphalis (tribe Gnaphalieae) is centred in Asia and is distinguished from the closely allied Antennaria and Helichrysum by its corymbose, scarious-bracted heads and its functionally dioecious breeding system; A. xylorhiza belongs to the woody-rooted, high-elevation Himalayan group. Its dense indumentum and persistent papery bracts are adaptations to cold, irradiation and desiccation in the alpine zone. Widespread and locally abundant, it is not considered threatened, although heavy summer grazing affects accessible populations.

Author: Sch.Bip. ex Hook.f. · Sources: POWO (Kew), GBIF, Flora of China, Annotated Checklist of the Flowering Plants of Nepal, Hooker Flora of British India (Asteraceae)

Asteraceae

Artemesia bhutanica

by G & P Jürgens

Griersson & Spring.

Bhutan2015 310 ergebnis
Bhutan Trongsa 1800m, October
Bhutan2015 311 ergebnis
Bhutan Trongsa 1800m, October

Description

Artemisia bhutanica is a perennial herbaceous species endemic or near-endemic to Bhutan, where it inhabits warm-temperate and subtropical forest margins, roadsides, and disturbed slopes at elevations of approximately 1,200–2,500 m. The species is recorded from the central valleys of Bhutan, including the Trongsa district at around 1,800 m, where it has been photographed flowering in October. With only nine occurrence records documented in GBIF, all confined to Bhutan, the species appears to have a highly restricted distribution and is one of the lesser-known members of this large, cosmopolitan genus in the eastern Himalayan region.
The plant forms erect to ascending, somewhat woody-based stems 40–100 cm tall, branching in the upper portion. The leaves are alternate, deeply pinnatisect to bipinnatisect, with narrowly linear to filiform segments, greyish-green above and more densely tomentose beneath with appressed T-shaped trichomes characteristic of the genus. The capitula are small, nodding, ovoid to subglobose, 2–3 mm in diameter, arranged in lax, paniculate synflorescences. Each capitulum contains few tubular florets; the outer florets are pistillate and the inner ones bisexual. The involucral bracts are imbricate, scarious-margined. Achenes are minute, obovoid, and glabrous.
Artemisia bhutanica was described by Grierson and Springate in the Flora of Bhutan and represents part of the considerable Artemisia diversity in the eastern Himalayas. Its taxonomic placement within the genus requires further molecular study, as morphological boundaries among eastern Himalayan Artemisia species can be unclear. Given its extremely limited documented range and low number of collections, A. bhutanica may warrant conservation assessment, particularly as road construction and land-use change continue to alter warm-temperate habitats in Bhutan's central valleys.

Author: Griersson & Spring. · Sources: POWO (Kew), GBIF, Grierson & Long: Flora of Bhutan Vol. 2 (2001)

Asteraceae

Artemisia austroyunnanensis

by GP Jürgens

Ling & Y.R.Ling

Bhutan2015 310
Bhutan Trongsa 2000m, October

Description

Artemisia austroyunnanensis Ling & Y.R.Ling is a Sino-Himalayan and South-East Asian wormwood distributed from south-western China (notably southern Yunnan, whence the epithet) through Bhutan, India, Myanmar, Laos, Vietnam, Thailand and Japan, with about 70 georeferenced occurrences in GBIF. It grows on dry slopes, forest margins, roadsides, riverbanks and disturbed ground in warm-temperate to lower montane country. The plant recorded at Trongsa at 2000 m in October is consistent with its occurrence in the mid-elevation valleys of the Bhutan Himalaya, flowering toward the close of the growing season.

It is an aromatic perennial herb or subshrub with erect, ribbed and branching stems clothed, like the foliage, in a fine grey or whitish tomentum. The leaves are alternate, once or twice pinnatifid into narrow, often lobed segments, with the lower surface more densely tomentose than the upper. The minute, nodding capitula are numerous and crowded in elongate, paniculate inflorescences; each ovoid to subglobose head has imbricate, woolly-margined phyllaries and is disciform, with outer female florets surrounding inner bisexual disc florets, all wind-pollinated and lacking rays. The fruit is a small, smooth, ellipsoid cypsela without a pappus.

Artemisia is among the largest genera of Asteraceae, and A. austroyunnanensis sits within the subgenus Artemisia of anemophilous, capitulum-clustered species. Its delimitation rests on indumentum, leaf dissection and capitulum structure within a difficult species complex. A regionally distributed plant of disturbed and seasonally dry habitats, it is of no conservation concern.

Author: Ling & Y.R.Ling · Sources: POWO (Kew), GBIF, Flora of China, Flora of Bhutan, Ling taxonomic studies of Artemisia

Asteraceae

Aster albescens

by GP Jürgens

(DC.) Wall. ex Hand.-Mazz.

Sikkim2003 379 G532
Sikkim Tista valley 3600m August
Garw 2005 38 ergebnis ergebnis
India Garhwal Valley of flowers 3200m, September

Description

Aster albescens is a variable shrubby aster widespread through the Himalaya and into China, Myanmar and East Asia, with 1,298 GBIF occurrences from China, Nepal, India, Bhutan, Japan, Korea and Myanmar. It grows on forest margins, open slopes, shrubberies and stream banks in the upper temperate to subalpine zone, broadly between about 2000 and 4000 m. The images derive from Sikkim's Tista valley at 3600 m in August and from the Valley of Flowers in Garhwal, India, at 3200 m in September, monsoon-watered slopes where this late-summer flowerer is a frequent component of the shrub layer.

Unusually for an aster, it is a woody shrub or subshrub up to about 1-2 m, with erect, branched, often whitish-tomentose stems. The leaves are alternate, ovate to lanceolate or elliptic, shortly petiolate, with serrate to entire margins and a characteristically pale, white-felted lower surface (whence albescens), the upper surface greener and thinly hairy. The capitula are numerous and small, arranged in dense terminal corymbose panicles; each head bears few short white, pinkish or pale purple ray florets around a disc of yellow florets. The cypselae are compressed and pubescent, with a pappus of reddish to brownish barbellate bristles.

Described as Diplopappus albescens by de Candolle and transferred to Aster by Wallich ex Handel-Mazzetti, the species is highly polymorphic and has spawned numerous infraspecific taxa across its range. Its shrubby habit places it among the woody Himalayan-Sino asters retained in Aster sensu lato. Common and ecologically resilient across a broad area, it is not considered threatened.

Author: (DC.) Wall. ex Hand.-Mazz. · Sources: POWO (Kew), GBIF, Flora of Bhutan, Flora of China, Flowers of the Himalaya (Polunin & Stainton), Asteraceae of the Himalaya

Asteraceae

Aster diplostephioides

by GP Jürgens

(DC.) Benth. ex C.B.Clarke

Bhutan2012 248
Bhutan Haar-Valley, 4000m September

Description

Aster diplostephioides is a striking alpine daisy of the high Himalaya and Tibetan borderlands, recorded by GBIF (322 occurrences) from China, India, Nepal and Bhutan. It inhabits open alpine meadows, grassy slopes, stony pastures and stream margins, typically between about 3000 and 4500 m. The plant illustrated was photographed in Bhutan's Haa (Haar) Valley at 4000 m in September, a high subalpine-to-alpine zone where short summers and intense insolation favour large-headed, single-flowered composites that bloom into the post-monsoon period.

It is a perennial herb arising from a stout, woody rhizome, producing erect, simple, glandular-hairy stems usually to 30-50 cm. The leaves are alternate, oblanceolate to spathulate, entire, somewhat clasping above, and clothed in spreading hairs. Each stem bears a single large, solitary capitulum subtended by herbaceous, often purple-tinged phyllaries in two or three rows. The numerous narrow ray florets are blue to violet-purple and conspicuously long, surrounding a central disc of yellow florets that turns orange-purple with age. The cypselae are flattened and hairy, bearing a pappus of unequal barbellate bristles.

The species was described as Heterochaeta diplostephioides by de Candolle and transferred to Aster by Bentham ex C.B. Clarke. Although molecular work has redistributed much of the former broad Aster, the high-elevation Himalayan asters such as this remain referable to Aster sensu stricto. Its conspicuous solitary heads make it a notable component of Himalayan alpine meadows. It is widespread within its montane range and not regarded as threatened, though alpine habitats face pressure from grazing intensification and climate warming.

Author: (DC.) Benth. ex C.B.Clarke · Sources: POWO (Kew), GBIF, Flora of Bhutan, Flora of China, Flowers of the Himalaya (Polunin & Stainton), Asteraceae of the Himalaya

Asteraceae

Aster falconeri

by G & P Jürgens

(C.B.Clarke) Hutch.

Nepal 99 93 ergebnis
West-Nepal Bhulbhule 3600m, June
Nepal 99 258 ergebnis
West-Nepal Loccu 3155m, June
Nepal 99 122 ergebnis
West-Nepal South of Rara-Lake 3500m, June

Description

Aster falconeri is a high-alpine herb of the north-western and western Himalaya, recorded from India, Pakistan and Nepal, with some 147 georeferenced collections documenting a relatively narrow but well-sampled range. It occupies open turf, moist alpine meadows, grazed grassland and the margins of seepages and snowmelt flushes, typically between about 3100 and 4000 m. The photographed populations from West Nepal—Bhulbhule at 3600 m, Loccu at 3155 m and the slopes south of Rara Lake at 3500 m, all flowering in June—are characteristic of its early-summer phenology, the plants emerging and blooming soon after the snowpack recedes.

It is a perennial forming low tufts from a short rootstock, the stems erect to ascending, simple, glandular-hairy and bearing reduced cauline leaves. Basal and lower leaves are oblanceolate to spathulate, narrowed to a winged petiole, entire or obscurely toothed and somewhat hairy. The solitary terminal capitula are radiate, subtended by an involucre of subequal, herbaceous, often purple-tinged phyllaries; the ray florets are ligulate and blue to violet, surrounding a disc of yellow tubular florets. The fruit is a compressed, hairy cypsela crowned by a pappus of barbellate bristles aiding wind dispersal.

First described as a member of Erigeron by C. B. Clarke and transferred by Hutchinson, the taxon is treated under the name Aster falconeri, regarded as the accepted name notwithstanding its synonym status in the source dataset. As with many Sino-Himalayan asters, generic limits between Aster and Erigeron have shifted repeatedly with morphological and molecular reappraisal. Though not formally assessed for conservation, its restriction to montane pastures exposes it to heavy grazing pressure across the western ranges.

Author: (C.B.Clarke) Hutch. · Sources: POWO (Kew), GBIF, Flora of Pakistan, Annotated Checklist of the Flowering Plants of Nepal, Grierson & Long Flora of Bhutan, Grierson's revision of Himalayan Aster (Asteraceae)