Tag:

Myanmar

Flowers of the Myanmar Himalaya — 604 species recorded from northern Myanmar (Burma) and the eastern Himalayan borderlands, a biodiversity hotspot linking the Himalaya to the mountains of Yunnan and Southeast Asia. The region’s forests and alpine slopes hold a rich flora of rhododendrons, orchids, balsams and gentians. Every page gives a botanical description, distribution, habitat and flowering time with the original photograph.

Ericaceae

Rhododendron vaccinioides

by GP Jürgens

Hook.

Bhutan2014 811
Bhutan Wangdi Mo-Chu 1600m, May
Nepal 97 108 P184
Nepal Dongen 2400m April
Nepal 1994 196 ergebnis
Ost-Nepal Mewa-Khola
2100m April

Description

Rhododendron vaccinioides is a small epiphytic rhododendron of the eastern Himalaya and adjacent China and Southeast Asia, recorded from China, India, Myanmar, Nepal, Bhutan, Vietnam and Japan, with some 245 GBIF occurrences. It grows as an epiphyte on mossy trunks and branches, and occasionally on rocks, in humid temperate broadleaved and conifer forest, generally between about 1600 and 3000 m. The documented populations span Wangdi Mo-Chu in Bhutan at 1600 m, Dongen in Nepal at 2400 m and the Mewa Khola in eastern Nepal at 2100 m, flowering in April and May in the cloud-bathed mid-montane belt.

It is a small, much-branched epiphytic shrub with slender, often pendent or sprawling branchlets bearing scattered scales (lepidote indumentum). The leaves are small, obovate to oblanceolate and somewhat fleshy, clustered toward the branch tips, vaccinium-like in aspect, hence the epithet, and lepidote beneath. The flowers are small, solitary or few at the branch ends, with a short-tubed, broadly funnelform to rotate, white to pinkish corolla that is sparsely scaly outside. The fruit is a small lepidote capsule splitting into valves to release numerous minute seeds adapted to wind dispersal among the canopy.

The species belongs to Rhododendron subgenus Rhododendron, section Vireya (the largely epiphytic, tropical-montane rhododendrons), of which it is one of the few representatives reaching the Himalaya. It is widespread and not threatened, but as an epiphyte dependent on humid, moss-laden old-growth forest it is sensitive to deforestation and to drying of the montane cloud belt.

Author: Hook. · Sources: POWO (Kew), GBIF, Flora of Bhutan, Flora of China, Argent monograph of Rhododendrons of Subgenus Vireya

Ericaceae

Rhododendron vaccinioides

by GP Jürgens

Hook. f.

Bhutan2014 43
Bhutan Trashigang 1300m, May
Bhutan2014 391
Bhutan Tumching-La 3000m, May
Bh2008 146 P243
Bhutan Chendebji, 2450m April

Description

Rhododendron vaccinioides Hook. f. is a small epiphytic rhododendron of the eastern Himalaya and adjacent montane Asia, ranging from Nepal and India through Bhutan to southern China, Myanmar and Vietnam, with around 245 GBIF occurrences. It grows characteristically as an epiphyte on mossy trunks and branches, and occasionally on rocks, in humid broadleaved and mixed evergreen forest, across a wide altitudinal span. The three Bhutan gatherings cited, from Trashigang at 1300 m and Chendebji at 2450 m flowering in April and May, and Tumching-La at 3000 m, illustrate its broad range from warm subtropical to cool temperate cloud forest where atmospheric moisture is high.

The species is a dwarf, much-branched epiphytic shrub with slender twigs clothed in scales and bearing small, obovate, leathery leaves whose lower surface is densely lepidote, resembling the foliage of Vaccinium as the epithet implies. The flowers are small, solitary or few together at the branch tips, with a short tubular to campanulate, white to pale pink corolla, the lobes spreading; the ovary and style are scaly. The fruit is a small lepidote capsule splitting septicidally to release numerous minute seeds adapted to wind dispersal in the canopy.

Rhododendron vaccinioides belongs to subgenus Rhododendron, section Vireya-allied scaly rhododendrons, an epiphytic lineage radiating through the wet forests of Asia. Its diminutive epiphytic habit is unusual among Himalayan rhododendrons. Widespread and locally frequent, it is of least conservation concern, though dependent on intact humid forest canopy.

Author: Hook. f. · Sources: POWO (Kew), GBIF, Flora of Bhutan, Flora of China, The Genus Rhododendron (Royal Botanic Garden Edinburgh), Cullen monograph of lepidote Rhododendron

Commelinaceae

Rhopalephora scaberrima

by GP Jürgens

(Blume) Faden

Bhutan2015 254
Bhutan Zhemgang 910m, October

Description

Rhopalephora scaberrima is a tropical to subtropical spiderwort widely distributed across South and Southeast Asia, recorded from India, Nepal, Bhutan, Bangladesh, Sri Lanka, Myanmar, Thailand, Malaysia, Indonesia, the Philippines, Vietnam, southern China and Taiwan. Its GBIF profile, with about 1,104 occurrences spanning these countries, confirms a broad Indomalayan range. The plant illustrated was photographed in southern Bhutan at Zhemgang at about 910 m in October. It is a species of warm, humid lowland and lower-montane settings, growing in forest understorey, along shaded streambanks, in moist grassy clearings and on disturbed roadsides, generally below about 1,500 m where high rainfall and warmth prevail.

It is a perennial, somewhat straggling herb with ascending to decumbent stems that root at the lower nodes, the surfaces and leaf-sheaths conspicuously scabrid-hispid, as the epithet indicates. The leaves are alternate, ovate-lanceolate to elliptic, with a closed sheathing base and an acuminate apex. The flowers are borne in small scorpioid cymes aggregated into a terminal panicle; each has three free sepals and three blue to violet petals, with the androecium showing the differentiation characteristic of the family, the fertile stamens separated from reduced staminodes. The fruit is a small loculicidal capsule, distinctively two-locular with the seeds arranged in two rows, a key feature separating Rhopalephora from related genera.

The genus Rhopalephora was segregated from the broadly defined Aneilema and Commelina on the basis of its bilocular capsule, biseriate seeds and stamen architecture, as clarified in Faden's revisionary work. As a common and adaptable plant of disturbed humid habitats, R. scaberrima is not threatened and serves as a useful indicator of warm, moist forest edge in the eastern Himalayan foothills.

Author: (Blume) Faden · Sources: POWO (Kew), Flora of China Vol. 24, Flora of Bhutan Vol. 3, GBIF, Faden: Commelinaceae (revisionary studies)

Anacardiaceae

Rhus chinensis

by G & P Jürgens

Mill.

Bhutan2012 535
Bhutan Wangdi 2600m, September
Bhutan2015 252
Bhutan Zhemgang 900m, October

Description

Rhus chinensis is a widespread deciduous shrub or small tree of East and South Asia, recorded across China, Taiwan, Japan, Korea, India, Vietnam, Thailand, Myanmar, Nepal, Bhutan and beyond, with more than ten thousand documented occurrences making it one of the most frequently collected sumacs in the region. It is a pioneer of open and disturbed ground, colonising forest margins, clearings, roadsides, scrub and abandoned cultivation from the lowlands into the montane zone. In Bhutan it ranges over a wide elevational band, as shown by collections near Wangdi at about 2600 m in September and at Zhemgang at 900 m in October, the autumn dates corresponding to fruiting.

The plant is a shrub to small tree with greyish bark and stout, often warted branchlets. The leaves are alternate and odd-pinnate, the rachis characteristically winged between the leaflets; the leaflets are ovate to oblong, coarsely serrate, acuminate, and tomentose beneath, turning red in autumn. The inflorescence is a large terminal pubescent panicle of numerous small whitish to greenish-yellow flowers, the plant functionally dioecious. The fruit is a small, flattened, reddish drupe covered with short hairs and a sour, salty surface bloom.

Rhus chinensis is the principal host of the Chinese sumac aphid Schlechtendalia chinensis, whose feeding induces large hollow leaf galls extraordinarily rich in tannins; these galls, known as Chinese galls or wubeizi, are harvested as a major commercial source of gallotannic acid and are valued in traditional medicine and dyeing. Wax from the fruit and the tannin-laden tissues have long been utilised economically. Hardy, fast-growing and broadly tolerant of disturbance, the species is abundant throughout its range and faces no conservation concern, functioning ecologically as an early-successional colonist.

Author: Mill. · Sources: POWO (Kew), GBIF, Flora of China Vol. 11, Flora of Bhutan Vol. 2, Flora of Nepal, Flora of Pakistan, Min & Barfod treatment of Anacardiaceae

Loxodinae

Rhynchoglossum obliquum

by GP Jürgens

Blume

Bhutan2012 983
Bhutan Trithangebe 1100m, October
Bhutan2012 984
Bhutan Trithangebe 1100m, October

Description

Rhynchoglossum obliquum is a widespread tropical Asian gesneriad ranging from India, Nepal, Bhutan and Bangladesh through Myanmar, Thailand, southern China, Taiwan and the Malesian region (Malaysia, Indonesia, Philippines, New Guinea, Timor) to Indochina, with more than 2200 GBIF occurrences. It is an annual herb of moist, shaded habitats on forest floors, damp rock faces, stream banks and roadside cuttings in the subtropical to tropical lowland and lower-montane zone. The plant recorded at Trithangebe in southern Bhutan at 1100 m, flowering in October at the close of the monsoon, typifies its occurrence on humid, shaded ground at low to mid elevations.

It is a soft, erect annual with succulent, often translucent stems. The leaves are markedly asymmetric (oblique) at the base, ovate to ovate-lanceolate, membranous and entire to obscurely toothed, the strong inequilaterality reflected in the epithet and frequently arranged so that the lamina appears one-sided. The flowers are borne in slender, secund, one-sided terminal racemes; each has a tubular, winged or keeled calyx and a small two-lipped blue to violet corolla with a short tube, the lower lip often marked white. There are two fertile stamens, and the superior ovary ripens into a small ellipsoid capsule enclosed by the persistent calyx, releasing numerous minute seeds.

The input places the species in the subtribe Loxonieae (here as Loxodinae) within Gesneriaceae, where Rhynchoglossum is distinguished by its oblique leaves and secund racemes. Its small bilabiate flowers are insect-pollinated and the dust-like seeds dispersed by rain-wash, aiding colonisation of disturbed shaded ground. As a widespread weedy annual it is of no conservation concern.

Author: Blume · Sources: POWO (Kew), GBIF, Flora of China Vol. 18, Flora of Bhutan Vol. 2, Flora of Nepal, The Gesneriaceae of the Old World (Rhynchoglossum)

Orchideaceae

Rhynchostylis retusa 

by GP Jürgens

(L.) Blume

Nepal 99 1024 ergebnis ergebnis
Nepal Kathmandu 1300m, July
Nepal 99 1025 ergebnis ergebnis
Nepal Kathmandu 1300m, July

Description

Rhynchostylis retusa (L.) Blume, the foxtail orchid, is an epiphyte widely distributed across tropical and subtropical Asia, with GBIF (617 occurrences) recording it from India, Nepal, Bhutan, China, Myanmar, Thailand, Laos, Vietnam, Cambodia, Sri Lanka, Bangladesh, Indonesia and the Philippines, plus many cultivated records. It grows on the trunks and branches of broadleaved trees in humid lowland and submontane forest, along forest edges and on roadside trees, flowering during the warm, wet summer. The photographed plant from the Kathmandu valley in Nepal at 1300 m in July reflects its typical occurrence in the warm-temperate to subtropical foothill belt of the Himalaya, where it flowers with the monsoon.

The species is a monopodial epiphyte with a short, stout stem clad in the persistent bases of distichous, fleshy, strap-shaped leaves that are channelled and conspicuously praemorse (retuse to unequally bilobed) at the apex, giving the epithet. Numerous thick, greyish aerial roots clasp the host bark. The inflorescence is a dense, pendent, cylindrical raceme of many small fragrant flowers, each white spotted and suffused with rose-purple or violet, with a spurred, entire to obscurely lobed labellum and a small column. The fruit is a pendent dehiscent capsule shedding dust-like wind-dispersed seeds.

Within the vandaceous alliance Rhynchostylis is a small genus of showy, scented epiphytes much esteemed in horticulture, and R. retusa holds particular cultural significance, being the state flower (kopou phul) of Assam. Like all orchids it is protected under CITES Appendix II. While still locally common, wild populations are reduced by habitat loss and unsustainable collection for the ornamental trade.

Author: (L.) Blume · Sources: POWO (Kew), GBIF, Flora of Nepal, The Orchids of the Himalaya (King & Pantling), Genera Orchidacearum (vandaceous orchids), CITES Appendix II

Grossulariaceae

Ribes himalense

by G & P Jürgens

Royle ex Decne.

Garh 07 113 ergebnis
India Garhwal Meggu Kichatti 3200m, June
Garh 07 114 ergebnis
India Garhwal Meggu Kichatti 3200m, June

Description

Ribes himalense Royle ex Decne. is a deciduous currant of the Himalaya and the mountains of western China. Its native range, reflected in GBIF records from India, Nepal, Pakistan, Bhutan, Afghanistan, China and Myanmar (with numerous European and North American records representing cultivated or naturalised material), spans the Himalaya into the Hengduan ranges. It inhabits subalpine forest, forest margins, streamsides and rocky slopes, typically between about 2,500 and 4,000 m. The plant from Meggu Kichatti in Garhwal at 3,200 m, fruiting in June, illustrates its occurrence in the upper temperate to subalpine belt.

The species is an erect, unarmed (spineless) shrub up to about 1–2 m tall with glabrous to glandular young branches. The alternate leaves are palmately three- to five-lobed, cordate at the base, with serrate lobes, glabrous or sparsely hairy. The flowers are borne in erect to spreading racemes; each flower has a shallowly saucer-shaped, greenish to purplish hypanthium, small reflexed sepals and minute petals, and is subtended by a bract. The fruit is a smooth, glabrous, red berry that is edible, ripening to a translucent crimson.

Ribes himalense belongs to the subgenus Ribes (the true unarmed currants) and is closely related to the European R. rubrum, with which it shares edible red fruit; it has been treated by some authors as a subspecies. The fruits are gathered locally and consumed, and the species is of interest as a wild relative of cultivated currants for breeding. It is widespread and of least concern.

Author: Royle ex Decne. · Sources: POWO (Kew), GBIF, Flora of China Vol. 8, Flora of Pakistan, Berger: A Taxonomic Review of Currants and Gooseberries

Grossulariaceae

Ribes luridum

by GP Jürgens

Hook. f. & Thomson

Bhutan2014 606
BhutanPhobjika 2810, May
Sikkim2011 76
Sikkim Tanghu 3900m, June

Description

Ribes luridum is a Sino-Himalayan currant distributed from south-western China through northern Myanmar, Nepal, northern India and Bhutan, with around 348 GBIF occurrences (the records from US, CA, PL and RU largely representing cultivated or herbarium material). It is a shrub of cool-temperate to subalpine forest, scrub margins and rocky slopes, typically between about 2800 and 4000 m. The collections from Phobjika in Bhutan at 2810 m in May and from Tanghu in Sikkim at 3900 m in June bracket this altitudinal range and its late-spring to early-summer flowering, on moist, humus-rich ground at and above the upper forest belt.

It is a deciduous, unarmed shrub with spreading branches and finely glandular young growth. The leaves are alternate, orbicular-cordate in outline and palmately three- to five-lobed, the lobes crenate-serrate, sparsely pubescent and often glandular beneath. The species is dioecious or functionally so, the small flowers borne in short, few-flowered, pendulous racemes; each has a saucer-shaped to shortly campanulate hypanthium and five small, dull purplish to lurid greenish-brown petals and sepals, the colouration alluded to by the epithet. The fruit is a small subglobose berry, glandular-bristly and reddish to dark purple when ripe.

Within Ribes the species sits in the section of unarmed currants and is distinguished by its dingy, glandular flowers and dioecious tendency. The pendulous berries are dispersed by frugivorous birds, an important agent in fragmented montane forest, while the inconspicuous flowers attract small bees and flies. The species is widespread across the eastern Himalaya and faces no recognised conservation threat.

Author: Hook. f. & Thomson · Sources: POWO (Kew), GBIF, Flora of China Vol. 8, Flora of Bhutan Vol. 1, Flora of Nepal, A Monograph of the Genus Ribes

Grossulariaceae

Ribes takare

by GP Jürgens

D. Don

Sikkim2011 365
Sikkim Lada-La 3900m, June
Bh2008 12 P20
Bhutan Thimphu-Chu, 2750m April
Sikkim2004 62 P110
Sikkim West-Bengal, Bersay 2700m April
Sikkim 2006 30 ergebnis
India West-Bengal Tonglu 3150m, May

Description

Ribes takare is a Himalayan currant distributed along the temperate and subalpine belt from Afghanistan through Pakistan, Nepal, India and Bhutan into southwestern China and Myanmar, with GBIF recording close to a thousand occurrences across this arc. It grows in moist temperate and upper montane forest, forest margins, shrubberies, ravines and rocky slopes, typically between about 2500 and 4000 m. The image caption documents this range well, with collections from the Lada La in Sikkim at 3900 m in June, the Thimphu Chu in Bhutan at 2750 m in April, Bersay in West Bengal at 2700 m in April, and Tonglu at 3150 m in May, showing both its broad altitudinal span and its spring-to-early-summer flowering as the snowline retreats.

It is a deciduous, spineless (unarmed) shrub up to about 2-3 m tall, with greyish, peeling bark and glandular-pubescent young shoots. The leaves are alternate, palmately three- to five-lobed with crenate-serrate margins and a cordate base, somewhat resembling a maple leaf and often glandular-hairy beneath. The flowers are borne in many-flowered, pendulous racemes; they are small and reddish to purplish-brown, with a saucer-shaped to shallowly campanulate hypanthium, five reflexed sepals, five smaller petals, five stamens and an inferior ovary. The fruit is a glandular-bristly, reddish to purplish-black berry that is edible but rather acidic and somewhat resinous.

The genus Ribes, sole genus of Grossulariaceae, is well represented in the Himalaya, where R. takare belongs to the unarmed, raceme-flowered black-currant alliance and has been treated under synonyms including R. acuminatum and R. glaciale-related names by various authors. Its pendent racemes of nectariferous flowers attract bees and other insects, while the berries are dispersed by birds and small mammals; the species forms an understorey component of moist Himalayan forest. It is widespread and common across its extensive range and is of no conservation concern, while its acidic fruits are gathered locally and its hardiness has drawn modest horticultural interest.

Author: D. Don · Sources: POWO (Kew), GBIF, Flora of China Vol. 8, Flora of Bhutan Vol. 1, Flora of Nepal, Flora of Pakistan, Berger's taxonomic review of Ribes

Asparagaceae

Rohdea nepalensis

by GP Jürgens

(Raf.) N.Tanaka

Nepal1992 155 P227 ergebnis
Nepal Simbua-Khola 2000m March 

Description

Rohdea nepalensis is a rhizomatous perennial herb of the Himalaya and adjacent montane Asia, documented by GBIF from China, India, Nepal, Bhutan, Myanmar, Bangladesh and Thailand across some 133 occurrences. It is a forest-floor species of shaded, moist broadleaved and evergreen forest, growing on humus-rich slopes, banks and rocky ground in the subtropical to warm-temperate zone. The photographed plant was recorded in Nepal in the Simbua-Khola at 2000 m in March, an early-season flowering in the cool, damp understorey characteristic of the species.

It forms clumps from a short, stout rhizome and produces basal, distichous, strap-shaped to oblanceolate leaves that are leathery, arching, prominently veined and tapered to a sheathing base. The inflorescence is a short, dense, sessile or shortly pedunculate spike borne near ground level, carrying numerous small, crowded flowers; each flower has a fleshy, urceolate to campanulate perianth of six connate tepals, usually pale yellow to greenish, with six included stamens and a superior ovary. The fruit is a one- to few-seeded berry that ripens red.

GBIF accepts the name, recombined by N. Tanaka, who expanded Rohdea to include species formerly placed in Tupistra and Campylandra; under that classification the plant appears in older literature under those genera. It is treated within an enlarged Asparagaceae (subfamily Nolinoideae, the former Convallariaceae/Ruscaceae). Several Rohdea are cultivated as evergreen ornamentals and used in traditional medicine. Widely distributed in suitable forest, R. nepalensis is of no conservation concern.

Author: (Raf.) N.Tanaka · Sources: POWO (Kew), GBIF, Flora of China, Flora of Nepal, Tanaka revision of Rohdea