Tag:

Nepal

Flowers of Nepal — 1313 wild plant species documented from the Nepalese Himalaya, from subtropical foothills to high alpine meadows above 4,000 m. Nepal’s dramatic elevation gradient makes it one of the richest botanical regions on Earth, home to rhododendrons, primulas, gentians and the famed blue poppies (Meconopsis). Every entry gives a botanical description, native distribution, habitat and flowering season, alongside the original field photograph and the location where it was taken.

Asteraceae

Leontopodium jacotianum

by GP Jürgens

Beauverd

Bhutan2012 57
Bhutan Shele-La, 4100m September
Bhutan2012 61
Bhutan Shele-La, 4100m September

Description

Leontopodium jacotianum is a high-alpine edelweiss of the Sino-Himalayan region, recorded from Nepal, China, India, Bhutan, Pakistan and Myanmar (NP, CN, IN, BT, PK, MM), with a modest 546 GBIF occurrences reflecting its restriction to lofty, difficult terrain. It inhabits open alpine meadows, stony slopes, screes and rocky crevices above the treeline, where snow lies long and the growing season is brief. The photographed plant grew on the Shele-La in Bhutan at about 4100 m in September, flowering at the close of the monsoon season on exposed alpine turf typical of the genus across the eastern Himalaya.

It is a tufted perennial herb arising from a stout, branched caudex, forming compact cushions or small clumps of several flowering stems 5–25 cm tall, the whole densely grey-white lanate-tomentose. The basal and cauline leaves are spathulate to linear-oblong, sessile, woolly on both surfaces. The diagnostic feature is the terminal star of leaf-like floral bracts radiating beyond the capitula, densely felted with white woolly hairs and forming the familiar edelweiss "flower". Within this stellate ruff sit several crowded capitula; the heads contain yellowish tubular florets, the plants typically subdioecious with outer female and central functionally male florets. The fruit is a small cypsela crowned by a pappus of barbellate to subplumose bristles.

The genus Leontopodium (tribe Gnaphalieae) reaches its greatest diversity in the Himalaya and the mountains of western China, where numerous species are differentiated by indumentum, bract shape and habit; L. jacotianum, described by Beauverd, belongs to this dense Asiatic radiation. The thick woolly investment is an adaptation to intense ultraviolet radiation, desiccation and cold at high elevation. Though not currently assessed as threatened, such narrowly alpine specialists are sensitive to climatic warming and the upslope contraction of suitable habitat.

Author: Beauverd · Sources: POWO (Kew), GBIF, Flora of China Vol. 20-21, Flora of Bhutan, Flora of Nepal, Leontopodium monograph (Handel-Mazzetti, revision of the genus)

Lamiaceae

Leonurus cardiaca

by G & P Jürgens

L.

Garh 07 310 ergebnis
India Garhwal Bebra 2450m, June

Description

Leonurus cardiaca, motherwort, is a perennial herb of Eurasian origin that has become widely naturalised, with a very large GBIF dataset of 79501 occurrences concentrated in Europe, Russia and temperate North America. It is a plant of disturbed, nutrient-rich ground, occurring on roadsides, hedgerows, waste places, field margins, fallows and around habitation. The plant recorded at Bebra in the Garhwal Himalaya at 2450 m in June represents an introduced or naturalised montane occurrence, where it grows on open, anthropogenically disturbed ground and flowers in early summer.

The species is an erect perennial reaching 0.5-1.2 m, with the quadrangular stems typical of Lamiaceae, often branched and somewhat pubescent. The leaves are opposite and decussate, long-petiolate, the lower palmately three- to five-lobed with coarsely toothed lobes, grading to narrower, fewer-lobed leaves above. The small flowers are borne in dense axillary verticillasters subtended by stiff, spinescent bracteoles; each has a five-toothed, rigid, sharply pointed calyx and a two-lipped pinkish to pale purple corolla, the upper lip densely white-villous and the lower three-lobed, enclosing four didynamous stamens. The fruit comprises four trigonous nutlets.

Leonurus belongs to the subfamily Lamioideae, and L. cardiaca has long been valued in herbal medicine as a cardiac and nervine tonic, the basis of both its scientific and common names. As a hardy, seed-prolific ruderal, it spreads readily in disturbed temperate habitats and is locally established well beyond its native Eurasian range; it is of no conservation concern and is sometimes regarded as a minor weed.

Author: L. · Sources: POWO (Kew), GBIF, Flora of China Vol. 17, Flora of Bhutan Vol. 2, Flora of Nepal, Flora of Pakistan, Krestovskaja's revision of Leonurus

Brassicaceae

Lepidostemon williamsii

by GP Jürgens

(H.Hara) Al-Shehbaz

BhutanPG2011 417
Bhutan Ludlow-Sherriff-Trail Warrtlang-La, 4400m June

Description

Lepidostemon williamsii is a rare high-alpine crucifer known only from Nepal and Bhutan, and is among the most sparsely documented species in this account, with merely about 12 GBIF records. It is confined to the upper alpine zone, growing on open screes, moraines, gravelly slopes and rock crevices near the limit of vascular plant life. The single cited collection, from the Ludlow-Sherriff-Trail at Warrtlang-La in Bhutan at about 4400 m in June, exemplifies its restriction to cold, exposed, well-drained mineral substrates where snow lies late and the growing season is extremely short.

The plant is a small, often cushion-forming or low tufted perennial herb adapted to severe alpine conditions, with a basal cluster of leaves that are entire to shallowly toothed or lobed. The flowers are typical of the family, with four sepals and four clawed petals arranged in a cross and arrayed in a short, congested raceme. The genus Lepidostemon is diagnosed by its distinctive stamens, whose filaments bear conspicuous scale-like or winged appendages, a feature reflected in the generic name. The fruit is a short silicula or silique characteristic of high-alpine Brassicaceae, splitting to release the seeds.

The species was originally described by H. Hara and transferred to Lepidostemon by Al-Shehbaz, whose revisions have clarified the circumscription of the small Sino-Himalayan genera of Brassicaceae. As a narrow-range endemic occupying a very specialised and harsh high-alpine niche with very few known populations, L. williamsii is inherently vulnerable, and its restricted distribution makes it a priority for monitoring against habitat disturbance and climatic change.

Author: (H.Hara) Al-Shehbaz · Sources: POWO (Kew), GBIF, Flora of Bhutan, Annotated Checklist of the Flowering Plants of Nepal, Al-Shehbaz's revision of Himalayan Brassicaceae

Trichosporeae

Leptoboea multiflora

by GP Jürgens

(C.B. Clarke) Benth. ex Gamble

BhutanPG2011 823 01 01
Bhutan Paro 3000m, July

Description

Leptoboea multiflora is a perennial herb of the eastern Himalaya and adjacent Southeast Asia, with 153 GBIF occurrences recorded from India, Nepal, Bhutan, China, Myanmar, Thailand, Vietnam and Cambodia. As a member of the Gesneriaceae, it grows in shaded, humid forest, typically on mossy rocks, boulders, banks and tree bases in damp, sheltered situations of the temperate broad-leaved zone, often between about 1500 and 3000 m. The plant photographed at Paro in Bhutan at 3000 m in July occupies the upper end of this range, flowering during the moist monsoon period when atmospheric humidity is high.

The species is a low, herbaceous to slightly succulent perennial with short stems and leaves that may be clustered or subopposite, the blades ovate to elliptic, crenate to serrate at the margins and frequently pubescent, with the asymmetric, anisophyllous tendency common in the family. The flowers are numerous, as the epithet indicates, borne in axillary cymes; each has a small five-lobed calyx and a tubular, somewhat bilabiate corolla, with the typically reduced fertile stamen number of the gesneriad tribe and a superior ovary. The fruit is a slender, elongate capsule dehiscing to release very numerous minute seeds.

The entry lists the family as Trichosporeae, which is in fact a tribe of Gesneriaceae rather than a family; the accurate placement is Gesneriaceae, subfamily Didymocarpoideae, tribe Trichosporeae, within which Leptoboea is a small Asian genus allied to Boea and Paraboea. As a humidity-dependent epilithic forest herb, it is sensitive to canopy disturbance, though it remains locally frequent in undisturbed montane forest.

Author: (C.B. Clarke) Benth. ex Gamble · Sources: POWO (Kew), GBIF, Flora of China Vol. 18, Flora of Bhutan Vol. 2, Flora of Nepal, Flora of Pakistan, Weber's classification of Gesneriaceae

Rubiaceae

Leptodermis stapfiana

by G & P Jürgens

H.J.P.Winkl

Bhutan2012 1036
Bhutan Trithangebe 900m, October

Description

Leptodermis stapfiana H.J.P.Winkl. is a shrub of the Rubiaceae distributed through the central and eastern Himalaya and into adjacent China. GBIF records it from Nepal, Bhutan, India and China, with 76 georeferenced occurrences. The species favours warm to cool temperate slopes, forest margins, scrub and roadside thickets, typically on well-drained, often rocky ground. The illustrated plant from Trithangebe in Bhutan at 900 m, flowering in October, reflects its preference for the lower montane belt, where it can form conspicuous patches of bloom late in the growing season after the main monsoon.

The plant is an erect to spreading, much-branched shrub with slender, greyish, often finely pubescent branchlets. The leaves are opposite, small, ovate to lanceolate, entire, shortly petiolate, and subtended by triangular, aristate stipules characteristic of the genus. Flowers are sessile and clustered in the leaf axils and at the branch tips; the corolla is narrowly funnel-shaped, white to pale lilac or pinkish, with five (rarely four) spreading lobes that are valvate and crisped-pubescent within the throat. The species is typically heterostylous, and the fruit is a small dry capsule splitting into valves to release angular seeds enclosed in a delicate membranous testa, the feature that gives Leptodermis its name.

Leptodermis is a taxonomically difficult Sino-Himalayan genus in which species are separated largely by leaf size, calyx-lobe number and indumentum, and L. stapfiana sits within this radiation centred on the eastern Himalaya and southwest China. It is not regarded as threatened, but as a hardy, late-flowering nectar source it contributes to montane pollinator networks and to the structural diversity of Himalayan scrub vegetation.

Author: H.J.P.Winkl · Sources: POWO (Kew), GBIF, Flora of China Vol. 19, Flora of Bhutan Vol. 2, Flora of Nepal, Deb & Rout: Studies in Indian Leptodermis (Rubiaceae)

Lamiaceae

Leucas decemdentata

by G & P Jürgens

(Willd.) Sm.

Sikkim 2006 260 ergebnis
India West-Bengal, Lingza 1300m, May

Description

Leucas decemdentata is one of the most widely distributed members of its genus, ranging across tropical and subtropical Asia and out into the western Pacific. GBIF records, totalling more than 1,600 occurrences, document its presence in India, Nepal, Pakistan and Myanmar through China, Taiwan and Japan to Thailand, the Philippines, Indonesia, Timor-Leste and Papua New Guinea, and onward to Australia, Fiji, Tonga, Samoa, the Cook Islands and French Polynesia, as well as Christmas Island. In the eastern Himalaya it is a plant of warm-temperate to subtropical situations, frequenting open hillsides, roadside banks, cultivated margins, fallow fields and other disturbed ground. The photographed plant from Lingza in West Bengal, India, at about 1300 m and flowering in May, is typical of its lower montane occurrence, where it behaves as a weedy ruderal of sunny, well-drained slopes.

The species is an erect or ascending annual to short-lived perennial herb, usually 30-100 cm tall, with quadrangular stems that are retrorsely hairy along the angles. Leaves are opposite and decussate, ovate to lanceolate, with a crenate-serrate margin, an acute apex and a cuneate to attenuate base, the lamina sparsely pubescent and borne on short petioles. The white flowers are crowded into dense, globose, axillary verticillasters subtended by linear bracts. Each flower has a tubular, ten-toothed calyx with subequal spinescent teeth and a strongly two-lipped corolla, the upper lip arched and densely white-villous, the lower lip three-lobed with a larger middle lobe; the four didynamous stamens lie beneath the hood. The fruit comprises four smooth, trigonous nutlets enclosed in the persistent calyx.

The ten conspicuous calyx teeth give the species its epithet decemdentata and are a useful diagnostic character within the large, taxonomically intricate genus Leucas. Long included with the white-flowered, capitate-inflorescenced "thumbergia group" of species, it has frequently been confused with L. ciliata, L. mollissima and other allies, and several Asian names have been treated as synonyms. Ecologically it is a pioneer of open, disturbed habitats and a minor melliferous plant visited by bees and other insects. It carries no conservation concern, being abundant and adaptable across an enormous geographic range, and like many Leucas species it is used locally in folk medicine for fevers and skin complaints.

Author: (Willd.) Sm. · Sources: POWO (Kew), GBIF, Flora of China Vol. 17, Flora of Bhutan Vol. 2, Flora of Nepal, Flora of Pakistan, Revision of the genus Leucas (Lamiaceae) by Ryding

Lamiaceae

Leucosceptrum canum

by G & P Jürgens

Sm.

Bhutan2015 595
Bhutan Mo-Chu 1900m, October
Bhutan2012 1307
Bhutan Mo-Chu 1900m, October
Nepal 95 429 P745 ergebnis
Nepal 95 428 P744 ergebnis
Nepal Trisuli 2700m October

Description

Leucosceptrum canum is a soft-wooded shrub or small tree of the Himalaya and southwestern China, distributed through China, Nepal, India, Bhutan and Myanmar and east to Indochina (CN, NP, IN, BT, MM, VN, TH, BD, LA, with further records), one of the more frequently collected species here with about 515 occurrences. It grows in moist temperate forest, on shaded slopes, ravine sides, forest margins and clearings, typically in the warm- to cool-temperate belt. The gatherings cited are from the Mo-Chu valley in Bhutan at 1900 m and from Trisuli in Nepal at 2700 m, both flowering in October, reflecting its autumn-to-early-winter blooming in the mid-montane forest zone.

The plant is a coarse shrub or small tree with stout, four-angled young branches densely clothed, like the leaf undersides and inflorescence, in a greyish (canous) tomentum from which the epithet derives. The leaves are large, opposite, elliptic to oblong-lanceolate, with a serrate margin, an acuminate apex and prominent venation. The flowers are crowded into a dense, terminal, cylindrical spike-like thyrse with conspicuous bracts; the calyx is tubular and five-toothed, and the white to pale yellow, two-lipped corolla has a short tube and exserted, declinate stamens. The fruit comprises four small nutlets (an eremocarp) enclosed in the persistent calyx.

Leucosceptrum is a small genus placed in the subfamily Lamioideae (or basal lineages) of the Lamiaceae and is of phylogenetic interest as part of an early-diverging Asian woody assemblage allied to Comanthosphace. The conspicuous, nectar-rich autumn inflorescences attract bees, butterflies and birds, and the species is notable for the production of defensive sesquiterpenes secreted by glandular trichomes. Widespread and common across the Himalaya into China, L. canum is of no conservation concern and is locally valued for fuel and as a bee plant.

Author: Sm. · Sources: POWO (Kew), GBIF, Flora of Bhutan, Flora of China, Labiatae of the Himalaya (Lamiaceae revision)

Caprifoliaceae

Leycesteria formosa

by G & P Jürgens

Wall.

Nepal 2009 106 ergebnis
Nepal Helambu Indrawati 2000, May
Nepal 2009 107 ergebnis
Nepal Helambu Indrawati 2000, May

Description

Leycesteria formosa is a Himalayan shrub native to the temperate forests of Nepal, India, Bhutan, Pakistan and southwestern China, where it grows in moist montane woodland, forest margins, ravines, streamsides and shrubby clearings, typically between about 1500 and 3000 m. The photographed plant from the Indrawati drainage in the Helambu region of Nepal, recorded in May, is characteristic of these cool, humid mid-elevation habitats. Far beyond its native range, however, the species is among the most frequently recorded plants in GBIF, with over 42,000 occurrences, the great majority from regions where it has become naturalised after cultivation as an ornamental, notably Great Britain, Ireland, France, the Netherlands, Belgium, Germany, Switzerland, Spain, Portugal, the United States, Australia and New Zealand, where it is regarded as an invasive woodland weed.

It is a deciduous, hollow-stemmed shrub 1-2.5 m tall, with erect, glaucous, bamboo-like green canes that are conspicuously jointed and unbranched or sparingly branched. The leaves are opposite, ovate to ovate-lanceolate, long-acuminate, entire to slightly toothed and often weakly cordate at the base, frequently with a few basal lobes. The inflorescences are pendent, interrupted spike-like racemes of whorled flowers subtended by showy, claret-purple, leaf-like bracts. Each funnel-shaped corolla is white flushed with purple-pink and five-lobed; the ovary is inferior. The fruit is a many-seeded berry that ripens from green through reddish-purple to a glossy deep maroon-black, soft and sweetish, much sought by birds, which disperse the abundant small seeds.

The genus Leycesteria, named for the Bengal magistrate William Leycester, comprises a small group of Sino-Himalayan shrubs traditionally placed in Caprifoliaceae (Lonicereae) near Lonicera. Bird-mediated seed dispersal underlies both the plant's ecological success and its problematic naturalisation abroad, where it colonises disturbed woodland and riparian thickets; in New Zealand in particular it is listed as an environmental weed. Within its native Himalayan range it is common and of no conservation concern, valued horticulturally for its colourful pendent inflorescences and ornamental berries, sometimes grown under the names "Himalayan honeysuckle" or "pheasant berry."

Author: Wall. · Sources: POWO (Kew), GBIF, Flora of China Vol. 19, Flora of Bhutan Vol. 2, Flora of Nepal, Flora of Pakistan, Airy Shaw's revision of Leycesteria

Caprifoliaceae

Leycesteria glaucophylla

by G & P Jürgens

(Hook. f. & Thomson) Hook.f.

Bhutan2014 312
Bhutan Tumching-La 2600m, May

Description

Leycesteria glaucophylla is an eastern Himalayan shrub distributed across India, China, Bhutan and Nepal and into Myanmar; GBIF documents about 113 occurrences, including a few cultivated records in Great Britain and New Zealand. It is a plant of temperate montane forest, occurring at forest margins, in shrubberies, along stream sides and in clearings, generally between about 2000 and 3000 m. The photographed individual grows in Bhutan at Tumching La at around 2600 m in May, a setting typical of this shade-tolerant honeysuckle relative within the cool, humid forests of the eastern Himalaya during the pre-monsoon period.

The species is an erect, deciduous shrub with hollow, often glaucous young stems. The leaves are opposite, ovate to lanceolate, acuminate, entire to slightly toothed, and conspicuously glaucous, particularly beneath, a feature reflected in the epithet glaucophylla. The flowers are borne in pendulous, bracteate terminal and axillary inflorescences subtended by leafy bracts; each has a tubular, funnel-shaped, white to pinkish corolla with five short lobes. The fruit is a many-seeded berry ripening to a dark purplish colour, attractive to birds which act as the principal seed dispersers.

Leycesteria glaucophylla (Hook.f. & Thomson) Hook.f. belongs to a small genus of Himalayan and Sino-Himalayan shrubs allied to Lonicera, placed in Caprifoliaceae; its showy bracts and pendulous spikes give the genus ornamental interest, and several species are cultivated. The species is restricted to the eastern Himalaya and adjacent ranges but is locally frequent in montane forest and not currently regarded as threatened. Its bird-dispersed berries aid colonisation of forest gaps and margins.

Author: (Hook. f. & Thomson) Hook.f. · Sources: POWO (Kew), GBIF, Flora of Bhutan, Flora of China, taxonomic revisions of Leycesteria (Caprifoliaceae)

Caprifoliaceae

Leycesteria gracilis

by GP Jürgens

(Kurz) Airy Shaw

Bh2008 326 P504
Bhutan Jakar, 3000m April

Description

Leycesteria gracilis is a Himalayan and Sino-Himalayan shrub, GBIF recording around 232 occurrences from China, India, Bhutan, Nepal and Myanmar, with scattered records from Japan. It grows in the understorey and at the margins of temperate broadleaved and mixed forest, in shrubberies, along forest tracks and on shaded, moist banks, generally at mid to upper montane elevations between about 2400 and 3300 m. The plant photographed from Jakar (Bumthang) in central Bhutan at 3000 m, in early flower in April, is characteristic of its preference for cool, humid montane forest where it leafs and blooms in spring ahead of the summer rains.

It is a slender, erect to arching deciduous shrub with hollow, often glaucous young stems, more delicate in habit than its relative L. formosa—hence "gracilis." The leaves are opposite, ovate-lanceolate to lanceolate, long-acuminate, entire to finely toothed and thin in texture. The flowers are borne in pendulous, interrupted spikes or whorled clusters subtended by leafy, sometimes purplish bracts; each has a small calyx and a tubular to funnel-shaped, five-lobed corolla in shades of white, pink or pale purple. The fruit is a small, several-seeded berry ripening dark purplish to nearly black.

Transferred by Airy Shaw from Kurz's original epithet, L. gracilis belongs to the small Himalayan genus Leycesteria, now placed in Caprifoliaceae in its broad modern sense. It is locally frequent and not regarded as threatened, though dependent on intact montane forest. Ecologically its tubular flowers attract insect and bird pollinators, while its fleshy dark berries are taken by birds that disperse the seed, making it a useful component of regenerating montane scrub.

Author: (Kurz) Airy Shaw · Sources: POWO (Kew), GBIF, Flora of Bhutan, Flora of China, Airy Shaw revision of Leycesteria (Caprifoliaceae)