Author

G & P Jürgens

Ranunculaceae

Eriocapitella vitifolia 

by G & P Jürgens

(Buch.-Ham. ex DC.) Nakai

Bhutan2012 1325
Bhutan Pemagatse 1900m, October
Bhutan2012 447
Bhutan Wangdi 2500m, September
Garw 2005 255 ergebnis scaled
Garw 2005 256 ergebnis scaled
India Garhwal Sitel 1850m, September

Description

Eriocapitella vitifolia (formerly Anemone vitifolia) is a robust perennial herb widespread across the Himalaya and East Asia, recorded from China, Nepal, India, Bhutan, Taiwan, Myanmar, Pakistan and beyond, with nearly 2000 georeferenced occurrences in GBIF reflecting both its native range and ornamental cultivation. It favours forest margins, open slopes, shrubberies and disturbed banks, mainly in the warm- to cool-temperate belt between about 1500 and 3000 m. The plants illustrated were photographed in Bhutan at Pemagatse (1900 m) and Wangdi (2500 m) and in the Indian Garhwal at Sitel (1850 m), flowering from September into October.

The species arises from a stout, woody rootstock and produces long-petiolate basal leaves with broadly cordate-ovate, vine-like (vitifolia, grape-leaved) blades that are shallowly three- to five-lobed, toothed, and densely white-tomentose beneath. Tall, branching flowering scapes bear cymose clusters of showy flowers, each with five to many petaloid sepals (true petals absent) that are white to pale pink, surrounding a central mass of numerous yellow stamens and a domed cluster of carpels. The fruiting head is globose and conspicuously woolly, the densely hairy achenes being adapted for wind dispersal, a feature reflected in the generic name.

Eriocapitella was segregated from Anemone (Ranunculaceae) on molecular and morphological grounds, comprising the woolly-fruited Asian anemones, and E. vitifolia is one of its most familiar Himalayan representatives. It is widely grown in temperate gardens and is an ancestral parent of the cultivated Japanese anemones, explaining scattered records in Europe, North America and southern Africa. Containing acrid ranunculaceous compounds, it is avoided by grazers and, common and adaptable, is of no conservation concern.

Author: (Buch.-Ham. ex DC.) Nakai · Sources: POWO (Kew), GBIF, Flora of China, Flora of Bhutan, Annotated Checklist of the Flowering Plants of Nepal, Hoot et al. Phylogeny of Anemone and Allies

Lamiaceae

Eriophyton tuberosum 

by G & P Jürgens

(Hedge) Ryding

Nepal 99 751 ergebnis
West-Nepal Poksundo-Lake 3650m, July

Description

Eriophyton tuberosum is a high-alpine Himalayan labiate with a narrow distribution, GBIF holding only about 46 occurrences from Nepal, India and China. It is a plant of the alpine and subnival zones, occurring on open turf, scree margins, moist rocky slopes and stream-side flushes generally between about 3300 and 4300 m, enduring intense insolation, short growing seasons and cold nights. The population photographed beside Phoksundo (Poksundo) Lake in West Nepal at 3650 m, flowering in July at the height of the monsoon growing season, is typical of its restricted occurrence in cool, moist, high-elevation meadow and rock habitats of the western and central Himalaya.

It is a low perennial herb arising from a swollen tuberous rootstock, as the epithet indicates, with short, ascending to decumbent stems. The leaves are opposite and decussate, ovate to rounded, crenate, and—like many members of this alpine assemblage—somewhat fleshy and often woolly or villous, an adaptation to the harsh climate. The flowers are borne in axillary whorls toward the stem apex; each has a tubular, toothed calyx and a two-lipped corolla in shades of pink, purple or violet, with four didynamous stamens beneath the upper lip. The fruit comprises four smooth nutlets in the persistent calyx.

Transferred by Ryding to Eriophyton from Hedge's original placement, this species reflects a modern recircumscription absorbing several small Himalayan genera (notably Alajja) into an expanded Eriophyton of woolly alpine mints. Its very low occurrence count and narrow high-altitude band make it a regional endemic of conservation interest, sensitive to grazing and alpine warming, and a nectar source for high-altitude bumblebees.

Author: (Hedge) Ryding · Sources: POWO (Kew), GBIF, Flora of China, Annotated Checklist of the Flowering Plants of Nepal, Ryding revision of Eriophyton (Lamiaceae)

Brassicaceae

Erysimum benthamii

by G & P Jürgens

Monnet

Nepal 2009 88 ergebnis
Nepal Helambu Bolgoau 2050, May

Description

Erysimum benthamii is a Himalayan crucifer recorded from China, Nepal, Bhutan and India, with over 230 georeferenced occurrences spanning the central and eastern Himalaya. It favours open, sunny situations on dry banks, stony slopes, forest clearings and disturbed ground in the temperate and subalpine zones. The plant illustrated from Helambu at Bolgaon (about 2050 m) in Nepal in May reflects its typical mid-elevation habitat, where it flowers in spring on well-drained, often rocky or gravelly soils along trails and open hillsides.

It is an erect biennial or short-lived perennial herb covered, like most members of the genus, in appressed medifixed (malpighiaceous) two- to several-branched hairs that give the foliage a greyish cast. The stem is simple or branched above and bears narrowly lanceolate to linear, entire or remotely denticulate leaves. The flowers are arranged in elongating terminal racemes; the four sepals are erect, and the four clawed petals are bright yellow to orange-yellow in the cruciform arrangement typical of Brassicaceae. The fruit is a long, slender, four-angled silique held more or less erect on a short pedicel, splitting at maturity to release two rows of small seeds.

Erysimum is a large Holarctic genus whose Himalayan representatives are taxonomically intricate, distinguished by trichome branching, fruit shape and indumentum density; E. benthamii is set apart by these vegetative and silique characters. The presence of an indeterminate-country record (coded ZZ) reflects imprecise georeferencing rather than any disjunct range. Widespread and tolerant of disturbance, the species is of least conservation concern.

Author: Monnet · Sources: POWO (Kew), GBIF, Flora of Nepal, Flora of China, Flora of Bhutan, Polatschek's revisions of Erysimum

Brassicaceae

Erysimum pachycarpum

by G & P Jürgens

Hook.f. & Thomson

Nepal 2009 88 ergebnis
Nepal Samthang 2410m, May

Description

Erysimum pachycarpum is a Himalayan wallflower distributed across Nepal, Bhutan, northern India, and extending into southwestern China (Tibet, Yunnan). GBIF documents 39 occurrences from these countries. The species inhabits dry rocky slopes, open scrubland, disturbed ground along trails, and stony riverbanks in the warm to cool temperate zone, at elevations from approximately 1,800 to 3,500 m. The photograph from Samthang in Nepal at 2,410 m, taken in May, shows the species at a mid-elevation site during its spring flowering period.
The plant is a biennial or short-lived perennial herb, 20–60 cm tall, with erect, usually unbranched stems covered in appressed, medifixed (malpighian) trichomes characteristic of the genus. Basal leaves are oblanceolate, 3–8 cm long, with entire to sinuate-dentate margins, forming a rosette that may wither at flowering; cauline leaves are narrowly lanceolate, sessile, progressively smaller upward. The inflorescence is a dense, terminal raceme that elongates considerably in fruit. Flowers are tetramerous with yellow petals, 8–12 mm long, spatulate, clawed. Stamens six, tetradynamous. The silique is stout and thick-walled — hence the epithet pachycarpum (thick-fruited) — erect to slightly spreading, 3–6 cm long, quadrangular in cross-section, densely covered with appressed trichomes. Seeds are oblong, narrowly winged.
Erysimum pachycarpum is distinguished from other Himalayan Erysimum species primarily by its robust, thickened siliques and relatively large flowers. The genus Erysimum is taxonomically challenging due to frequent hybridisation and subtle morphological boundaries between species, and molecular studies are ongoing to resolve relationships within the Himalayan clade. The species is not currently assessed as threatened and appears locally common in suitable open, dry habitats across its range.

Author: Hook.f. & Thomson · Sources: POWO (Kew), GBIF, Flora of China Vol. 8, Grierson & Long: Flora of Bhutan Vol. 1

Brassicaceae

Erysimum robustum

by G & P Jürgens

D.Don

Nepal 99 423 ergebnis
Nepal Talphi 2830m, July
Nepal 99 72 ergebnis
Nepal Chauta-valley, 2800m, July

Description

Erysimum robustum is a Himalayan wallflower distributed through Nepal, northern India and SW China (Xizang and Yunnan), as reflected in the GBIF country records (NP, IN, CN) and some 89 occurrences. The plants illustrated here were photographed in western Nepal at Talphi (c. 2,830 m) and in the Chauta valley (c. 2,800 m) in July. It is a montane to subalpine species favouring open, well-drained sites: stony pastures, dry slopes, roadside banks and the margins of cultivation, generally between 2,500 and 3,800 m, where it forms conspicuous yellow-flowered stands during the summer monsoon. The GBIF backbone treats the name as a synonym, but it is here regarded as the accepted name Erysimum robustum following regional floristic usage.

It is a biennial or short-lived perennial herb with an erect, robust, often branched stem clothed, like the rest of the plant, in appressed two- to several-rayed malpighiaceous hairs that give a greyish cast. The leaves are linear to oblanceolate, entire to remotely denticulate, the lower ones petiolate and the upper sessile. The inflorescence is an elongating ebracteate raceme of bright yellow flowers with four clawed petals in the cruciferous pattern and tetradynamous stamens. The fruit is a long, narrow, four-angled siliqua, somewhat torulose and ascending on a stout pedicel, containing a single row of oblong seeds in each locule.

The genus Erysimum is taxonomically difficult owing to extensive variation in indumentum and habit, and several Himalayan entities have been variously lumped or split; trichome branching patterns and siliqua characters remain the most reliable diagnostic features. As a widespread, weedy-tolerant species of disturbed montane ground, E. robustum is not of conservation concern and can act as an early colonist of bare slopes.

Author: D.Don · Sources: POWO (Kew), Flora of China Vol. 8, Flora of Nepal, GBIF, Polatschek: Revision der Gattung Erysimum

Phrymaceae

Erythranthe nepalensis

by G & P Jürgens

(Benth.) G.L.Nesom

Bhutan2014 855
Bhutan Thimphu 2600m, May

Description

Erythranthe nepalensis is a small wetland herb of temperate East and South Asia, recorded from Japan, China, Korea, Taiwan, Nepal, Bhutan, India, Myanmar, Pakistan and Bhutan, with around 2199 georeferenced occurrences in GBIF. It grows in damp, open ground: stream banks, ditch margins, wet meadows, seepages, rice-field bunds and moist rock crevices from the foothills into the temperate montane zone. The photographed plant from Thimphu in Bhutan at 2600 m in May is characteristic of such cool, moist Himalayan habitats, where it flowers in spring along watercourses and on dripping banks.

It is a slender annual or short-lived perennial herb with weak, decumbent to ascending, often rooting stems forming low mats. The leaves are opposite, small, ovate to broadly ovate, palmately veined, with a few shallow teeth and short petioles. The flowers are solitary in the leaf axils on slender pedicels; the calyx is tubular-campanulate and angled-ribbed, with short triangular lobes, and the corolla is bilabiate and yellow, the throat marked with reddish-brown to purplish spots and the lower lip prominent. There are four didynamous stamens, and the fruit is an oblong capsule enclosed by the persistent ribbed calyx, releasing numerous tiny seeds.

Formerly placed in Mimulus (as Mimulus tenellus var. nepalensis), the species was transferred to Erythranthe by G.L. Nesom following the molecular recircumscription that divided the old Mimulus among several genera and moved them from Scrophulariaceae to Phrymaceae. The spotted, bilabiate yellow flowers are adapted to bee pollination, and the plant's affinity for moist, disturbed ground allows ready local colonisation. Common and widespread across its range, it is of no particular conservation concern.

Author: (Benth.) G.L.Nesom · Sources: POWO (Kew), GBIF, Flora of China Vol. 18, Flora of Bhutan Vol. 2, Flora of Nepal, Taxonomy of Erythranthe (Phrymaceae) (Nesom)

Celastraceae

Euonymus frigidus

by G & P Jürgens

Wall.

Sikkim 2006 184 ergebnis
India West-Bengal, Samadan 2300m, May
BhutanPG2011 82 ergebnis scaled
Bhutan Menchuang 2850m, June

Description

Euonymus frigidus is a Himalayan spindle tree distributed from central Nepal through northeast India, Sikkim, Bhutan, and northern Myanmar to southwest China (Yunnan, Sichuan, Xizang). GBIF records over 1,570 occurrences, with the majority from China and the eastern Himalayas. The species inhabits cool-temperate and subalpine broadleaved and mixed forests, often growing along forest margins, stream gullies, and shaded slopes at elevations of 1,800–3,200 m. Collections from Samadan in West Bengal at 2,300 m and from Menchuang in Bhutan at 2,850 m illustrate its typical occurrence in the mid-elevation broadleaved forest zone of the eastern Himalayan slopes.
Euonymus frigidus is a large deciduous shrub or small tree, reaching 3–8 m in height, with smooth, greyish-green bark on younger branches. Leaves are opposite, elliptic to oblong-lanceolate, 6–14 cm long, acuminate at the apex, cuneate at the base, with finely crenate-serrate margins, glabrous on both surfaces, and turning reddish-purple in autumn. The inflorescence is an axillary, dichasial cyme bearing 7–15 flowers on slender peduncles. Flowers are tetramerous, small, 6–8 mm in diameter, with greenish-yellow to purplish petals and a conspicuous four-lobed disc. The fruit is a deeply four-lobed capsule, pink to crimson at maturity, splitting to reveal four seeds each enclosed in a bright orange aril — a striking ornamental feature.
The specific epithet 'frigidus' alludes to the cold-temperate habitats in which this species predominantly occurs. Euonymus frigidus is distinguished from the closely related E. hamiltonianus by its generally larger leaves, more deeply lobed capsules, and preference for higher elevations. The conspicuous arillate fruits are an important food source for frugivorous birds, including minivets and laughingthrushes, which serve as effective dispersal agents. The species is occasionally cultivated as an ornamental in European and North American gardens for its vivid autumn colour and decorative fruit.

Author: Wall. · Sources: POWO (Kew), GBIF, Flora of China Vol. 11, Flora of Bhutan Vol. 2, Grierson & Long: Flora of Bhutan (1991)

Euphorbiaceae

Euphorbia pseudosikkimensis 

by G & P Jürgens

(Hurus. & Tanaka) Radcl.-Sm.

Sikkim 2006 323 ergebnis
India Sikkim Ringi-Chu 2250m, May
Sikkim 2006 464 ergebnis
India Sikkim Ringi-Chu 2250m, May
Sikkim 2006 324 ergebnis
India Sikkim Ringi-Chu 2250m, May

Description

Euphorbia pseudosikkimensis is a robust perennial spurge native to the eastern Himalayas, occurring in Nepal, Sikkim, Bhutan, and adjacent parts of China. It inhabits moist forests, forest clearings, and stream banks at 1,800–3,000 m.

The plant forms tall, erect stems up to 1.5 m high, with whorled, oblong-lanceolate leaves that are dark green above and paler beneath. The inflorescence is a large, showy cyathium with conspicuous bright yellow-green glands, giving the impression of striking yellow flowers in spring. Like all Euphorbia species, it exudes a milky white latex when stems or leaves are broken.

In traditional medicine of the region, the latex has been used cautiously as a purgative, though it is irritant and toxic in larger quantities. The species is sometimes confused with E. sikkimensis, from which it differs in leaf shape and gland morphology.

Author: (Hurus. & Tanaka) Radcl.-Sm. · Sources: POWO (Kew), Flora of Bhutan, eFlora of China

Euphorbiaceae

Euphorbia sharmae

by G & P Jürgens

U.C.Bhattach.

Garh 07 272 ergebnis
India Garhwal Belak Khal 2900m, June

Description

Euphorbia sharmae is a poorly known spurge apparently endemic to the western Himalaya of India, where it is recorded only from the Garhwal Himalaya of Uttarakhand. With very few documented occurrences, it appears to be a narrow regional endemic of temperate montane habitats, growing on open grassy slopes, forest clearings and rocky banks. The plant photographed at Belak Khal in Garhwal at 2,900 m in June occupies the cool temperate belt, flowering early in the warm season; the scarcity of records underscores how localised and infrequently collected the species is.

Like other members of the genus, the plant is a herbaceous perennial exuding milky white latex when damaged. It produces erect to ascending stems from a perennating rootstock, bearing alternate, entire, oblong to lanceolate cauline leaves and a terminal umbel-like arrangement of branches subtended by a whorl of leaf-like ray-leaves. The reproductive units are cyathia, the specialised pseudanthia diagnostic of Euphorbia, each comprising a cup-shaped involucre bearing crescent-shaped nectar glands and enclosing reduced unisexual flowers around a central stalked ovary. The fruit is a three-lobed schizocarpic capsule splitting into mericarps to release the seeds.

Euphorbia is among the largest and most cosmopolitan of angiosperm genera, and its Himalayan herbaceous representatives belong to the diverse subgenus Esula. Euphorbia sharmae, described by U. C. Bhattacharyya, is one of several narrowly distributed Indian endemics whose limits remain incompletely understood owing to sparse collection. Its restricted known range and the absence of population data place it among taxa of potential conservation concern, warranting targeted survey to clarify its distribution and status.

Author: U.C.Bhattach. · Sources: POWO (Kew), Flora of India (BSI), Flora of Uttar Pradesh, GBIF, Radcliffe-Smith: Euphorbia systematics

Euphorbiaceae

Euphorbia sikkimensis

by G & P Jürgens

Boiss.

Nepal 2001 302 ergebnis ergebnis
Nepal (Gorkha) Lambra Kharka 3100m, May
Nepal 2001 301 ergebnis
Nepal (Gorkha) Lambra Kharka 3100m, May
Nepal 97 86 P146
Nepal Tankhu 2100m April

Description

Euphorbia sikkimensis is a robust herbaceous spurge distributed across the eastern and central Himalaya, from central Nepal through Sikkim, Bhutan, and northeast India into southern China (Xizang, Yunnan), with peripheral records from Myanmar, Laos, and Vietnam. It occurs in subalpine meadows, forest clearings, scrub margins, and open hillsides at elevations of 1,800–3,500 m. The species has been documented from the Gorkha district of Nepal at Lambra Kharka at 3,100 m in May and near Tankhu at 2,100 m in April, indicating a broad elevational amplitude and early-season phenology at lower altitudes.
The plant is a stout, erect perennial herb, 30–80 cm tall, with thick, fleshy rootstock and unbranched or sparingly branched stems that exude copious white latex when cut. Leaves are alternate, sessile, linear-lanceolate to oblanceolate, 8–15 cm long and 1–2.5 cm wide, with a prominent pale midrib, entire margins, and a glaucous undersurface. The inflorescence is a terminal, compound cyathium typical of the genus, with a broad umbel of 4–8 primary rays, each ray subtended by obovate, yellowish-green floral bracts. The cyathia bear crescent-shaped glands with short horns. The fruit is a smooth, trilocular capsule approximately 6–8 mm in diameter, explosively dehiscent at maturity, with carunculate seeds.
Euphorbia sikkimensis was described by Boissier from Sikkim collections and belongs to Euphorbia subgenus Esula, a predominantly Eurasian clade of herbaceous spurges. It is notable for its striking appearance in early spring, when the bright red or pink-tinged new stems contrast with the yellowish-green inflorescences. The species has gained horticultural recognition in temperate gardens and is increasingly cultivated as an ornamental perennial in Europe. Ecologically, its copious latex deters herbivory, and it plays a role in early-season pollinator support in montane grassland ecosystems.

Author: Boiss. · Sources: POWO (Kew), GBIF, Flora of China Vol. 11, Annotated Checklist of the Flowering Plants of Nepal