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Myanmar

Flowers of the Myanmar Himalaya — 604 species recorded from northern Myanmar (Burma) and the eastern Himalayan borderlands, a biodiversity hotspot linking the Himalaya to the mountains of Yunnan and Southeast Asia. The region’s forests and alpine slopes hold a rich flora of rhododendrons, orchids, balsams and gentians. Every page gives a botanical description, distribution, habitat and flowering time with the original photograph.

Ericaceae

Lyonia villosa

by G & P Jürgens

(Wall. ex C.B. Clarke) Hand.-Mazz.

Lyonia villosa ergebnis scaled
Bhutan Wangdi, 2400m June

Description

Lyonia villosa (Wall. ex C.B.Clarke) Hand.-Mazz. is a widely distributed Sino-Himalayan shrub recorded from China, Nepal, Bhutan, India, Myanmar, Vietnam and Japan, with more than 1300 georeferenced occurrences in GBIF, reflecting its broad range across the eastern Himalaya and southern China. It grows in temperate forest, forest margins, thickets and open rocky hillsides, commonly between about 1500 and 3500 m. The plant photographed near Wangdi at 2400 m in June is characteristic of its mid-elevation habitat, where it forms part of the understorey and scrub of mixed broadleaved and conifer woodland.

It is a deciduous shrub or small tree with villous to tomentose young branchlets, the indumentum giving rise to the epithet. The leaves are alternate, ovate to elliptic, entire, with a pubescent surface and prominent venation. The white, urceolate (urn-shaped) flowers are borne in axillary racemes or fascicles along the branches; each corolla is shortly five-lobed at the contracted mouth, with included stamens whose anthers bear awns, and a superior ovary maturing into a woody, dehiscent, five-valved capsule. The capsule has thickened sutures, a diagnostic feature of the genus, and releases numerous small seeds.

Lyonia belongs to the tribe Lyonieae of the Ericaceae and is allied to Pieris and Agarista; the thickened capsule sutures and awned anthers separate it from these relatives. Several varieties have been recognised within L. villosa on the basis of indumentum density and leaf shape. As in many Ericaceae the foliage is reported to be toxic to livestock. The species is common and widespread and is not considered threatened.

Author: (Wall. ex C.B. Clarke) Hand.-Mazz. · Sources: POWO (Kew), GBIF, Flora of China, Flora of Bhutan, Judd revision of Lyonia (Ericaceae)

Primulaceae

Lysimachia congestiflora

by G & P Jürgens

Hemsl.

Bhutan2014 782
Bhutan Wangdi Mo-Chu 1200m, May

Description

Lysimachia congestiflora is a wide-ranging species native across China, Taiwan, the Himalaya (Nepal, north-east India, Bhutan) and into Myanmar, Vietnam, Laos, Thailand and Japan; its very large total of 1844 georeferenced records includes numerous naturalised and cultivated points in the United States, Brazil, Ecuador, Mexico, Australia and Europe, reflecting its popularity as a creeping ground-cover and basket plant. It grows in moist, shaded to open ground on grassy banks, streamsides, ditches, field margins and damp roadside slopes. The Bhutan collection at Wangdi along the Mo-Chu at about 1200 m typifies its preference for warm, humid lower-montane habitats.

It is a low, mat-forming perennial herb with slender, decumbent to creeping stems that root at the lower nodes and are sparsely villous. The leaves are opposite, ovate to broadly elliptic, often gland-dotted and minutely hairy. The flowers are crowded in dense terminal clusters (hence congestiflora), each on a short pedicel; the corolla is yellow, rotate to broadly campanulate, five-lobed and frequently marked with reddish spots or a darker throat. There are five stamens, and the fruit is a small globose capsule enclosed by the persistent calyx.

Within Primulaceae the genus Lysimachia (sometimes segregated as Lysimachiopsis or placed in subgenus Idiophyton) comprises loosestrifes characterised by rotate corollas and capsular fruit. The clustered yellow flowers attract a range of generalist insect pollinators. Exceptionally adaptable and frequently weedy in cultivation, the species is abundant throughout its native range and of no conservation concern; its capacity to naturalise has even made it a minor garden escape in several regions.

Author: Hemsl. · Sources: POWO (Kew), GBIF, Flora of China, Flora of Bhutan, Flora of Nepal, Hu & Kelso Primulaceae monographic treatments

Primulaceae

Lysimachia evalvis

by G & P Jürgens

Wall.

Sikkim 2006 282 ergebnis
Sikkim 2006 281 ergebnis
India West-Bengal, Chaana 1650m, May

Description

Lysimachia evalvis is a Sino-Himalayan herb distributed along the moist temperate and subtropical belt of the eastern Himalaya and adjacent Indo-Burmese ranges. GBIF records (88 occurrences) span India, Nepal, Myanmar, Bhutan, Bangladesh and China, with scattered cultivated or vouchered material noted from the United States. It favours shaded, humid forest margins, stream banks and seepage slopes within broadleaved woodland. The species was photographed at Chaana in West Bengal, India, at about 1650 m in May, in the lower montane forests where it flowers with the onset of the pre-monsoon rains.

It is a perennial, somewhat fleshy herb with slender erect to ascending stems arising from a short rhizomatous base. The leaves are alternate to subopposite, ovate to lanceolate, entire, acuminate at the apex and often gland-dotted, borne on short petioles. The flowers are solitary or few in the upper leaf axils, with a deeply five-lobed calyx and a yellow, rotate to campanulate corolla divided into five lobes; the five stamens are inserted on the corolla tube. The fruit is a small globose capsule; as the epithet evalvis suggests, dehiscence is reduced or irregular rather than by the typical valves, a diagnostic feature within the genus.

Lysimachia (Primulaceae, formerly segregated in Myrsinaceae alliance) is a large, near-cosmopolitan genus well represented in the Himalaya. L. evalvis belongs to the predominantly Asian element of the genus, its distribution reflecting the warm, perhumid forests of the eastern Himalayan front. No formal conservation assessment is widely published; like many forest-floor herbs of this region, its persistence depends on intact humid montane forest, making habitat conversion the principal threat to local populations.

Author: Wall. · Sources: POWO (Kew), GBIF, Flora of China, Flora of Bhutan, Annotated Checklist of the Flowering Plants of Nepal, Hu's monograph of Lysimachia (Primulaceae)

Primulaceae

Lysimachia lobelioides

by GP Jürgens

Wall.

Bh09 395 P626
Bhutan Mo-Chu, 1350m June
Bh09 396 P628
Bhutan Mo-Chu, 1350m June

Description

Lysimachia lobelioides is a slender herb ranging widely through the Himalaya and Southeast Asia, recorded from China, India, Bhutan, Nepal, Myanmar, Thailand, Laos, Vietnam, Taiwan and New Caledonia, with 647 occurrences logged in GBIF. It favours moist, shaded and disturbed places at low to middle elevations: stream banks, ditches, damp grassy ground, field margins and open forest, broadly between 300 and 2500 m. The Bhutanese collection illustrated, from the Mo-Chu valley at 1350 m in June, typifies its occurrence in warm subtropical river-valley habitats during the early monsoon, when it flowers freely on damp, fertile soils.

It is an annual or short-lived perennial with weak, erect to decumbent, often quadrangular stems 10-40 cm long, glabrous and sometimes rooting at the lower nodes. The leaves are alternate, ovate to lanceolate, thin in texture, shortly petiolate and frequently dotted with dark glandular punctations. The small flowers are borne singly in the upper leaf axils on filiform pedicels; the corolla is white to pale pink, deeply five-lobed and shorter than or about equalling the lanceolate calyx lobes, with five stamens. The fruit is a small globose capsule dehiscing by valves and releasing numerous minute angular seeds.

Lysimachia (Primulaceae, formerly Myrsinaceae in part) is a large, near-cosmopolitan genus best developed in eastern Asia; L. lobelioides belongs to a group of small-flowered, axillary-flowered annuals. Its wide distribution, weedy tendency and tolerance of disturbed wet ground make it ecologically adaptable and of no conservation concern. The disjunct New Caledonian records are biogeographically notable, pointing to long-distance dispersal of this colonising species across the Indo-Pacific.

Author: Wall. · Sources: POWO (Kew), GBIF, Flora of China, Flora of Bhutan, Revision of the Genus Lysimachia (Primulaceae)

Primulaceae

Lysimachia prolifera

by GP Jürgens

Klatt

Nepal 2001 255 ergebnis ergebnis
Nepal (Gorkha) Niuli Kharka 2700m, May
Nepal 2001 256 ergebnis ergebnis
Nepal (Gorkha) Niuli Kharka 2700m, May

Description

Lysimachia prolifera is a Sino-Himalayan species distributed across the central and eastern Himalaya and southwestern China, with verified occurrences in Nepal, India, Bhutan, China, Myanmar and Pakistan; outlying records from Japan and the United States among the roughly 194 collections almost certainly reflect cultivated or misdetermined material. It is a plant of moist temperate to subalpine habitats, favouring shaded forest margins, streamsides, mossy banks and damp grassy clearings. The photographed population at Niuli Kharka in Gorkha, Nepal, at about 2700 m and flowering in May, is typical of its mid-montane band where snowmelt and monsoon moisture keep the substrate reliably humid.

The species is a perennial herb with slender, often decumbent to ascending stems arising from a creeping rhizomatous base. The leaves are simple, opposite or sometimes whorled, ovate to lanceolate, entire-margined and frequently gland-dotted, narrowing to a short petiole or sessile base. The flowers are borne singly in the leaf axils or in lax leafy racemes; the corolla is rotate and five-lobed, typically yellow, the lobes spreading and often marked with darker glands. Stamens are five, opposite the corolla lobes and shortly connate at the base, the ovary superior with a single style. The fruit is a small, globose, dehiscent capsule containing numerous minute angular seeds.

Within Primulaceae, Lysimachia is a large, taxonomically intricate genus now circumscribed broadly, and species such as L. prolifera are distinguished by subtle differences in leaf arrangement, gland distribution and inflorescence form that have historically caused confusion. Ecologically it contributes to the herbaceous understorey of Himalayan forests and is visited by generalist pollinating insects. No specific conservation concern is recorded for this widespread montane species, though, like many Himalayan herbs, it is locally sensitive to forest clearance, trampling and grazing along trekking corridors.

Author: Klatt · Sources: POWO (Kew), GBIF, Flora of Nepal, Flora of China, Flora of Bhutan, Hu's monograph of Lysimachia (Primulaceae)

Didymocarpinae

Lysionotus pubescens

by G & P Jürgens

C.B. Clarke

Bhutan2012 996
Bhutan Thritangebe 1000m, October

Description

Lysionotus pubescens is an epiphytic or lithophytic gesneriad of the humid subtropical to lower montane forests of the eastern Himalaya and Indo-Burma, recorded from China, India, Bhutan, Myanmar, Vietnam and Sri Lanka. Its 75 georeferenced occurrences trace a belt of warm, moist evergreen forest where it grows perched on mossy tree trunks, branches and shaded rock faces. The plant photographed at Thritangebe in Bhutan at 1000 m in October reflects its preference for the warm, low-elevation forest zone, flowering in the post-monsoon period when humidity remains high beneath the canopy.

The plant is a small woody-based perennial with pendent or ascending stems bearing leaves in whorls of three or more, or sometimes opposite. The leaves are leathery, elliptic to oblong, with a toothed (serrate to crenate) margin, and as the epithet indicates the herbage is finely pubescent. The flowers are borne in short axillary cymes; the corolla is tubular and distinctly two-lipped (bilabiate), white to pale pink or pale violet, often with yellow or purple markings in the throat, with two fertile stamens. The diagnostic fruit is a long, slender, linear capsule that dehisces to release numerous minute seeds bearing fine appendages.

Lysionotus belongs to subtribe Didymocarpinae of the Gesneriaceae, the family group cited here, and is distinguished within it by its long siliqua-like capsule and frequently whorled leaves. The genus is centred on the Sino-Himalayan and Indo-Chinese region and is well represented in China. As an epiphyte dependent on intact humid forest, L. pubescens is locally common where suitable habitat persists but is sensitive to forest clearance.

Author: C.B. Clarke · Sources: POWO (Kew), GBIF, Flora of China, Flora of Bhutan, Gesneriaceae of the Old World monograph (Weber), Flowers of the Himalaya (Polunin & Stainton)

Euphorbiaceae

Macaranga denticulata

by G & P Jürgens

Müll.Arg.

Bhutan2015 199
Bhutan, Zhemgang 910m, October
Bhutan2015 557
Bhutan, Punaka 1240m, October
Bhutan2015 556
Bhutan, Punaka 1240m, October

Description

Macaranga denticulata is a fast-growing pioneer tree with a wide tropical and subtropical Asian distribution, its 1451 georeferenced occurrences spanning China, India, Nepal, Bhutan, Bangladesh, Myanmar, Thailand, Vietnam, Laos, Cambodia, Malaysia, Indonesia, the Philippines and beyond. It is a characteristic colonist of disturbed ground, forest gaps, road cuttings, secondary forest and abandoned swidden plots in warm, humid foothill country. The records from Zhemgang at 910 m and Punakha at 1240 m in Bhutan in October place it firmly in the low-elevation subtropical zone where it regenerates rapidly after disturbance and forms early-successional thickets and woodland.

The plant is a small to medium evergreen tree with smooth bark and somewhat whorled, spreading branches. The leaves are large, alternate, peltate or subpeltate, broadly ovate to triangular with a denticulate margin reflected in the epithet, palmately veined, and often bearing glands and pale tomentum beneath. As a member of the Euphorbiaceae it is dioecious, the small apetalous flowers borne in branched axillary inflorescences subtended by toothed or glandular bracts; male and female flowers occur on separate trees. The fruit is a small two-lobed capsule, the seeds often associated with a fleshy aril attractive to birds and ants.

Macaranga is a large palaeotropical genus famous for its myrmecophytic (ant-associated) species and its ecological role as a pioneer. M. denticulata is a non-myrmecophytic but ecologically important coloniser, valued for rapid biomass accumulation, soil stabilisation and as a nurse species in fallows. Its enormous range, abundance and affinity for disturbed habitats make it among the most secure and widespread trees in its family across tropical Asia.

Author: Müll.Arg. · Sources: POWO (Kew), GBIF, Flora of China, Flora of Bhutan, Revision of Macaranga (Euphorbiaceae) (Whitmore and successors), Trees of tropical Asia

Lauraceae

Machilus kurzii

by G & P Jürgens

King ex Hook.f.

Sikkim2004 379 P656 ergebnis
Sikkim Lachung-valley 3200 m, May
Sikkim2004 380 P657 ergebnis
Sikkim Lachung-valley 3200 m, May

Description

Machilus kurzii is an evergreen laurel of the eastern Himalaya and Southeast Asia, distributed from NE India and Bhutan through Myanmar, Thailand, Cambodia and Vietnam into SW China, as documented by its roughly 135 GBIF occurrences across IN, BT, CN, MM, TH, KH, VN and KR. The tree photographed grows in the Lachung valley of northern Sikkim at about 3,200 m in May. It is a species of humid temperate and subtropical evergreen broadleaf forest, occurring on moist slopes and in ravines often dominated by other Lauraceae, Fagaceae and rhododendrons, where high rainfall and persistent cloud sustain the laurophyllous canopy across a broad altitudinal range.

It is an evergreen tree with smooth bark and glabrous to thinly pubescent twigs. The leaves are alternate, coriaceous, elliptic to oblanceolate, entire, with a pinnate venation of fine, looping secondary veins and an obtuse to shortly acuminate apex, often glaucous beneath. The flowers are small and bisexual, borne in subterminal or axillary panicles; they show the trimerous laurel plan with six tepals in two whorls, fertile stamens in several whorls including a third whorl with paired basal glands, and inward-facing anthers dehiscing by valves. The fruit is a globose to ellipsoid berry seated on a persistent, somewhat enlarged perianth base or cupule, characteristic of the genus.

Machilus, often treated within or close to Persea, is an Asian genus of the tribe Laureae whose delimitation has long been debated; molecular and floral evidence continues to refine the boundary between Machilus and Persea. The species is an integral element of mid-montane evergreen forest, contributing to canopy structure and providing fleshy fruits for birds. It is not currently considered threatened, though it depends on the maintenance of undisturbed humid broadleaf forest.

Author: King ex Hook.f. · Sources: POWO (Kew), Flora of China Vol. 7, Flora of Bhutan Vol. 1, GBIF, Kostermans: Lauraceae studies

Magnoliaceae

Magnolia campbellii

by GP Jürgens

Hook.f. & Thomson

Nepal 97 118 P204
Nepal Mewa-Khola 2470m April
Nepal 97 119 G206 2
Nepal Mewa-Khola 2470m April
Nepal1992 149 P216 ergebnis
Nepal Simbua-Khola 2500m March 

Description

Magnolia campbellii, Campbell's magnolia, is a large deciduous tree of the eastern Himalaya, native from Nepal through Sikkim and Bhutan into NE India, Myanmar and SW China; the additional GBIF codes (GB, IE, US, NZ, JP, DE and others) among its roughly 807 occurrences reflect its renown as a cultivated ornamental. The plants photographed grow in eastern Nepal in the Mewa-Khola (c. 2,470 m, April) and Simbua-Khola (c. 2,500 m, March) valleys, flowering before leaf flush in early spring. In the wild it inhabits temperate broadleaf and mixed forest on humid, well-drained slopes, typically between 2,100 and 3,000 m, often as an emergent tree among oak, rhododendron and laurel.

It is a tree reaching 20–30 m with a stout trunk, grey bark and stout, often pubescent young twigs marked by ringed stipular scars. The leaves are alternate, elliptic to obovate, entire, glaucous and finely hairy beneath. The large, solitary, precocious flowers are among the most spectacular of the genus, cup-and-saucer in form, 15–25 cm across, with numerous tepals ranging from deep rose-pink to white; as in all Magnoliaceae the receptacle bears spirally arranged stamens below an elongate gynoecium of free carpels. The fruit is an aggregate of follicles forming a cylindrical, often twisted etaerio that splits to expose red, arillate seeds suspended on slender funicular threads.

Magnolia campbellii belongs to subgenus Yulania, the precocious-flowering Asian magnolias, and its early bloom on bare branches is a defining feature exploited in horticulture, where numerous selections and hybrids exist. Though widely grown, wild populations face pressure from logging and forest conversion, and the species is regarded as of conservation interest across parts of its native range, making intact mid-montane Himalayan forest important to its persistence.

Author: Hook.f. & Thomson · Sources: POWO (Kew), Flora of China Vol. 7, Flora of Bhutan Vol. 1, GBIF, Hunt (ed.): Magnolias and their Allies

Magnoliaceae

Magnolia hodgsonii

by G & P Jürgens

(Hook.f. & Thomson) H.Keng

Sikkim 2006 492 ergebnis
India West-Bengal, Chaana 1700m, May

Description

Magnolia hodgsonii is an evergreen tree of the eastern Himalaya and adjacent Southeast Asia, distributed through northeast India, Nepal, Bhutan, southern China (Yunnan, Xizang), Myanmar, Thailand, Vietnam and into Malesia. It is a component of warm, humid subtropical to lower montane broad-leaved evergreen forest, favouring sheltered valleys, ravines and moist hill slopes on deep soils. It generally occurs from the lowlands to around 1,800 m; the tree photographed at Chaana in the Darjeeling hills of West Bengal at 1,700 m in May exemplifies its upper subtropical forest habitat, flowering at the close of the dry season.

The plant is a medium-sized to large tree with smooth grey bark and stout twigs. The leaves are large, alternate, often clustered toward the branch tips, obovate to elliptic-oblong, coriaceous, glabrous and entire, with a prominent venation. The solitary, terminal flowers are large and fragrant, with creamy-white to pale yellow tepals arranged in several whorls; the numerous stamens and the apocarpous gynoecium of free carpels are borne on an elongate floral axis. The fruit is an ellipsoid to ovoid aggregate of follicles which split to expose seeds enveloped in a fleshy red to orange sarcotesta, dispersed by birds.

The species, with basionym attributed to Hooker f. and Thomson and recombined by H. Keng, was formerly placed in the segregate genus Talauma, now subsumed within a broadly circumscribed Magnolia. As an archaic, beetle-pollinated member of one of the most basal angiosperm lineages, it is of considerable evolutionary interest. Valued for its timber and ornamental flowers, it is subject to selective logging and forest clearance, and warm-temperate populations on the forest fringe merit monitoring where habitat loss is ongoing.

Author: (Hook.f. & Thomson) H.Keng · Sources: POWO (Kew), Flora of China Vol. 7, Flora of Bhutan Vol. 1, GBIF, Figlar & Nooteboom: Magnoliaceae notes