Asteraceae

Asteraceae

Ajania myriantha

Asteraceae

Anaphalis busua

Asteraceae

Tricholepis elongata 

Asteraceae

Artemisia austroyunnanensis

Asteraceae

Asteraceae

Inula grandiflora 

by G & P Jürgens

Willd.

Garw 2005 26 ergebnis ergebnis
India Garhwal 2950m, September
Garw 2005 27 ergebnis ergebnis
India Garhwal 2950m September

Description

Inula grandiflora Willd. is a stout perennial composite of the western and central Himalaya, ranging from Pakistan and northern India through Nepal, with related records extending into the wider Irano-Turanian and Caucasian region (Turkey, Georgia, Armenia). It occupies open subalpine and alpine slopes, grassy meadows, forest clearings and rocky banks in the upper-temperate to alpine belt. The photographed plants from Garhwal in the Indian Himalaya at 2950 m, flowering in September, exemplify its preference for sunny, well-drained montane grassland where it flowers late in the season as the monsoon eases, frequently forming conspicuous stands on open hillsides.

The plant produces erect, leafy, often somewhat hairy stems from a woody rootstock. The alternate leaves are oblong to elliptic-lanceolate, sessile and frequently half-clasping the stem, with finely toothed margins and a roughened or pubescent surface. The inflorescence is a large, solitary or few-headed terminal capitulum (the epithet grandiflora referring to its size), subtended by overlapping herbaceous involucral bracts; the head bears numerous narrow, spreading golden-yellow ray florets surrounding a central disc of tubular florets. The fruit is a ribbed, cylindrical cypsela crowned by a pappus of fine bristles that aids wind dispersal.

Within the genus Inula (Asteraceae, tribe Inuleae) this is one of the showiest Himalayan species, valued horticulturally and used locally in traditional medicine. Its large, nectar- and pollen-rich heads attract bees, butterflies and other insects, making it an important late-season forage plant. Though not globally threatened and locally common, its alpine grassland habitats are subject to grazing and climate-driven upslope vegetation shifts.

Author: Willd. · Sources: POWO (Kew), GBIF, Flora of Pakistan, Annotated Checklist of the Flowering Plants of Nepal, Flora of China (Asteraceae), Compositae of the Himalaya

Asteraceae

Inula hookeri

by GP Jürgens

C.B.Clarke

Bhutan2012 181
Bhutan Haar-Valley, 3200m September
Bhutan2012 846
Bhutan Nangsephal, 2700m October
Bhutan2012 848
Bhutan Nangsephal, 2700m October

Description

Inula hookeri is a showy yellow-flowered composite of the eastern Himalaya, native to China, Bhutan, Nepal, India and Myanmar, and frequently cultivated in temperate gardens, which accounts for British, Irish, North American and Antipodean records; GBIF lists about 383 occurrences. It grows in subalpine meadows, open forest, shrubby slopes and grassy clearings, mostly between about 2400 and 3600 m. The photographed plants are from the Haar valley in Bhutan at 3200 m in September and from Nangsephal at 2700 m in October, flowering through the late monsoon into autumn.

The plant is an erect, hairy perennial herb arising from a short rhizome, with leafy, softly pubescent stems. The leaves are alternate, oblong-lanceolate to elliptic, acuminate, finely toothed and clasping or sessile at the base, hispid on both surfaces. The capitula are large and solitary or few at the stem tips, radiate, with numerous very narrow, thread-like, pale to greenish-yellow ray florets spreading in a fringed circle around a disc of deeper yellow tubular florets; the involucre is hemispherical with linear, leafy, spreading phyllaries. The fruit is a small ribbed cypsela bearing a pappus of a single row of barbellate bristles.

Inula belongs to the tribe Inuleae, and I. hookeri, named for J. D. Hooker, is distinctive for its softly spidery, narrow-rayed heads that open from woolly buds. Valued in horticulture for its long flowering and attractiveness to pollinators, it is common in the wild and not considered threatened.

Author: C.B.Clarke · Sources: POWO (Kew), GBIF, Flora of Bhutan, Flora of China, Annotated Checklist of the Flowering Plants of Nepal, Systematics of Inula and allied genera (Inuleae)

Asteraceae

Jacobaea raphanifolia

by G & P Jürgens

(Wall. ex DC.) B.Nord.

Sikkim2003 444 G617
Sikkim Chopta Valley 3900m August
Nepal 95 130 P222 ergebnis
Nepal south of Rupien La 3600m October
Bhutan2015 379
Bhutan Tumching-La 4000m, October

Description

Jacobaea raphanifolia is a perennial herb of the high Himalaya and adjoining mountains of central Asia, recorded by GBIF (some 202 occurrences) from Nepal, India, China, Bhutan, Afghanistan, Pakistan and Myanmar, with scattered records reaching Kenya and South Africa that probably reflect cultivated or naturalised material. It is a plant of the upper subalpine and alpine zones, favouring moist meadows, streamsides, open scrub and damp rocky slopes. The species was photographed across a broad altitudinal band, in the Chopta Valley of Sikkim at about 3900 m in August, south of the Rupien La in Nepal at 3600 m in October, and on the Tumching La in Bhutan near 4000 m, a range centred between roughly 3500 and 4200 m.

It forms an erect, somewhat stout herb arising from a short, fibrous-rooted rhizome, with simple or branched leafy stems. The lower leaves are lyrate-pinnatifid to runcinate with a large terminal lobe, recalling a radish (raphanifolia), the margins toothed and the surfaces sparsely hairy; cauline leaves become smaller, sessile and often auriculate upwards. The capitula are radiate, several to many in a loose terminal corymb, subtended by a single row of narrow green phyllaries with a few short calyculate bracts. Ray florets are yellow, female and ligulate; the disc florets yellow, tubular and bisexual. The fruit is a ribbed, cylindrical cypsela crowned by a pappus of fine white capillary bristles.

The species belongs to a complex of montane ragworts segregated from Senecio into Jacobaea on molecular and morphological grounds. As a wide-ranging meadow herb of the alpine pastures it is locally common and of no conservation concern, though like many Senecioneae it contains pyrrolizidine alkaloids and is generally avoided by grazing stock, a trait that may favour its persistence in heavily pastured high valleys.

Author: (Wall. ex DC.) B.Nord. · Sources: POWO (Kew), GBIF, Flora of Bhutan, Flora of China, Annotated Checklist of the Flowering Plants of Nepal, Nordenstam's revision of Jacobaea and the Senecioneae

Asteraceae

Lactuca dolichophylla 

by G & P Jürgens

Kitam.

Garw 2005 68 ergebnis ergebnis
India Garhwal Valley of flowers 3200m, September

Description

Lactuca dolichophylla is a Himalayan and Central Asian perennial herb distributed through the Indian Himalaya, Pakistan, Nepal, Bhutan, southern China, Afghanistan, Russia and Myanmar, documented from a modest number of georeferenced collections. It is a plant of the subalpine and alpine zone, growing on open grassy slopes, herb-rich meadows, forest clearings and stony banks. The photographed material is from the Valley of Flowers in Garhwal, India, at about 3200 m in September, a celebrated high-montane meadow habitat where the species flowers in the late post-monsoon season.

The plant is an erect perennial herb with a leafy, often glaucous stem exuding milky latex when cut. The leaves are elongate and narrow, as the epithet 'dolichophylla' (long-leaved) implies, lanceolate to linear-lanceolate, entire to runcinate-lobed, with the upper leaves sessile and often clasping. The capitula are arranged in a lax paniculate-corymbose synflorescence; each head is ligulate, with an imbricate cylindrical involucre and several blue to bluish-purple or violet ray florets, a colour characteristic of the Himalayan blue-flowered lettuces. The fruit is a flattened, ribbed achene (cypsela) narrowed into a beak and crowned by a pappus of simple white capillary bristles.

Lactuca belongs to the tribe Cichorieae of Asteraceae, the dandelion-and-lettuce alliance with milky latex and ligulate florets. The blue-flowered Himalayan species, sometimes segregated under Cicerbita or Mulgedium, form a taxonomically intricate group, and this taxon was described by Kitamura. Wind-dispersed by its pappus, L. dolichophylla is a characteristic component of moist alpine meadows and is not regarded as threatened.

Author: Kitam. · Sources: POWO (Kew), GBIF, Flora of China, Flora of Pakistan, Kitamura's studies of Himalayan Cichorieae (Compositae)

Asteraceae

Laphangium affine

by GP Jürgens

(D.Don) Tzvelev

Sikkim2004 102 P173
India West-Bengal, Utaray 2000m April
Sikkim 2006 186 ergebnis
India West-Bengal, Samadan 2400m, May

Description

Laphangium affine is a widespread East and South Asian cudweed recorded from China, Japan, India, Taiwan, Vietnam, Nepal, Pakistan, Thailand, Georgia and Turkey, the roughly 99 georeferenced occurrences reflecting a broad continental range that extends westward into the Caucasus and Anatolia. It occurs from lowland to mid-montane elevations, the images documenting plants in West Bengal, India, at Utaray about 2000 m in April and Samadan about 2400 m in May. A plant of open, disturbed and seasonally moist ground, it colonises fields, fallows, roadsides, terrace banks, streamsides and weedy grassland, behaving as a fast-growing ruderal of cultivated and human-modified landscapes.

It is an erect annual or short-lived herb with simple or branched stems clothed, like the foliage, in a soft greyish-white woolly tomentum. The leaves are alternate, sessile, narrowly oblanceolate to linear-spathulate, entire-margined and densely lanate on both surfaces, the lower often withered at flowering. The numerous small capitula are aggregated into dense terminal and axillary glomerules; each head bears straw-coloured to yellowish, dry, scarious involucral bracts and contains many slender female florets surrounding a few central bisexual ones. The cypselae are minute and crowned by a caducous pappus of barbellate bristles, adapted for wind dispersal.

Under GBIF this name is treated as a synonym, with Laphangium affine itself standing as the accepted name; the taxon has been variously placed in Gnaphalium and Pseudognaphalium, reflecting the unsettled generic limits of the woolly cudweeds in tribe Gnaphalieae. Originally described by D. Don and transferred by Tzvelev, it is a successful weed across much of Asia, valued in some regions for medicinal and edible use. Given its wide range and ruderal ecology it is of no conservation concern.

Author: (D.Don) Tzvelev · Sources: POWO (Kew), GBIF, Flora of China, Flora of Pakistan, Gnaphalieae generic revision (Asteraceae)

Asteraceae

Leontopodium brachyactis

by G & P Jürgens

Gand.

Sikkim2003 428 P595
Sikkim Chopta Valley 4100m August
Garw 2005 158 ergebnis
India Garhwal valley of flowers 3550m, October

Description

Leontopodium brachyactis is a high-alpine edelweiss of the Sino-Himalayan and Central Asian uplands, recorded from India, Pakistan, Nepal and China, with scattered occurrences extending west into Kyrgyzstan, Tajikistan, Afghanistan and Russia; the c. 92 georeferenced collections trace a distribution centred on the inner Himalaya and Tibetan margins. It inhabits open alpine turf, stony slopes, moraines and grazed meadows above the treeline, typically between 3500 and 4500 m. The species was photographed in the Sikkim Chopta Valley at 4100 m in August and in the Garhwal Valley of Flowers at 3550 m in October, sites representative of its preference for cold, well-drained, seasonally snow-covered ground.

It is a low, tufted, woolly perennial herb arising from a branched woody caudex, the stems erect to ascending and densely white-lanate. The leaves are linear to oblanceolate, entire, grey-green above and clothed beneath in a felt of appressed hairs that reflects intense montane irradiance. The capitula are small, several to many, aggregated into a dense terminal glomerule and subtended by a stellate ruff of leaf-like, densely tomentose floral bracts forming the characteristic edelweiss "star." The heads are heterogamous and disciform, with whitish corollas; the fruit is a small ribbed cypsela bearing a pappus of barbellate bristles.

The genus Leontopodium (tribe Gnaphalieae) is taxonomically intricate, its species delimited by indumentum, bract arrangement and capitulum structure, and L. brachyactis belongs to the compact-headed Himalayan element. Its dense white tomentum is a classic adaptation to alpine cold, desiccating winds and ultraviolet exposure. Though locally frequent across a wide range and not presently considered threatened, populations in accessible valleys face pressure from heavy summer grazing and increasing tourism in the Himalayan flower meadows.

Author: Gand. · Sources: POWO (Kew), GBIF, Flora of China, Flowers of the Himalaya (Polunin & Stainton), Handel-Mazzetti revision of Leontopodium

Asteraceae

Leontopodium himalayanum

by GP Jürgens

DC.

BhutanPG2011 507
Bhutan Ludlow-Sherriff-Trail Shashuna, 4000m June
Nepal 99 528 ergebnis
West-Nepal Churta-Khola 3800m, June

Description

Leontopodium himalayanum is the characteristic Himalayan edelweiss, distributed across China, Nepal, India, Bhutan, Pakistan and Myanmar (CN, NP, IN, BT, PK, MM), with about 437 GBIF occurrences mapping a broad but strictly montane range along the chain. It occupies alpine and subalpine meadows, open grassy and stony slopes, screes and rocky banks, generally above the forest limit. The photographed plants were recorded on the Ludlow-Sherriff Trail at Shashuna in Bhutan at 4000 m and at the Churta-Khola in West Nepal at 3800 m, both in June, flowering early in the high-mountain season on exposed turf.

It is a tufted perennial herb growing from a short branched rhizome, producing several erect or ascending flowering stems 5–30 cm tall, the herbage greyish with a dense appressed white tomentum. The leaves are linear, linear-lanceolate or oblong-spathulate, sessile, woolly especially beneath. The inflorescence is crowned by a spreading rosette of lanceolate floral bracts much more densely white-lanate than the foliage, forming the radiating star that distinguishes the edelweiss. Several small subglobose capitula are clustered at the centre of this ruff; the heads bear yellowish tubular florets and are functionally subdioecious. The fruit is a minute cypsela tipped with a pappus of slender, barbellate to subplumose bristles facilitating wind dispersal.

Within Leontopodium (tribe Gnaphalieae) the Himalayan species form a closely allied group differentiated chiefly by indumentum density and bract proportions, and L. himalayanum (described by DC.) is the most widespread representative across the range. Its conspicuous woolly star is a classic alpine adaptation buffering the reproductive organs against ultraviolet exposure, cold and water loss. The species is common and not regarded as threatened, although picking and collection have locally reduced populations near trekking routes.

Author: DC. · Sources: POWO (Kew), GBIF, Flora of China Vol. 20-21, Flora of Bhutan, Flora of Nepal, Leontopodium monograph (Handel-Mazzetti, revision of the genus)

Asteraceae

Leontopodium jacotianum

by GP Jürgens

Beauverd

Bhutan2012 57
Bhutan Shele-La, 4100m September
Bhutan2012 61
Bhutan Shele-La, 4100m September

Description

Leontopodium jacotianum is a high-alpine edelweiss of the Sino-Himalayan region, recorded from Nepal, China, India, Bhutan, Pakistan and Myanmar (NP, CN, IN, BT, PK, MM), with a modest 546 GBIF occurrences reflecting its restriction to lofty, difficult terrain. It inhabits open alpine meadows, stony slopes, screes and rocky crevices above the treeline, where snow lies long and the growing season is brief. The photographed plant grew on the Shele-La in Bhutan at about 4100 m in September, flowering at the close of the monsoon season on exposed alpine turf typical of the genus across the eastern Himalaya.

It is a tufted perennial herb arising from a stout, branched caudex, forming compact cushions or small clumps of several flowering stems 5–25 cm tall, the whole densely grey-white lanate-tomentose. The basal and cauline leaves are spathulate to linear-oblong, sessile, woolly on both surfaces. The diagnostic feature is the terminal star of leaf-like floral bracts radiating beyond the capitula, densely felted with white woolly hairs and forming the familiar edelweiss "flower". Within this stellate ruff sit several crowded capitula; the heads contain yellowish tubular florets, the plants typically subdioecious with outer female and central functionally male florets. The fruit is a small cypsela crowned by a pappus of barbellate to subplumose bristles.

The genus Leontopodium (tribe Gnaphalieae) reaches its greatest diversity in the Himalaya and the mountains of western China, where numerous species are differentiated by indumentum, bract shape and habit; L. jacotianum, described by Beauverd, belongs to this dense Asiatic radiation. The thick woolly investment is an adaptation to intense ultraviolet radiation, desiccation and cold at high elevation. Though not currently assessed as threatened, such narrowly alpine specialists are sensitive to climatic warming and the upslope contraction of suitable habitat.

Author: Beauverd · Sources: POWO (Kew), GBIF, Flora of China Vol. 20-21, Flora of Bhutan, Flora of Nepal, Leontopodium monograph (Handel-Mazzetti, revision of the genus)

Asteraceae

Leontopodium nivale subsp. alpinum

by G & P Jürgens

(Cass.) Greuter

Sikkim2003 325 P462 ergebnis
Sikkim Samiti lake 4000m, July

Description

Leontopodium nivale subsp. alpinum, the classic alpine edelweiss, is a high-mountain perennial of the Asteraceae with a remarkably broad circum-montane distribution. GBIF records (over 20,000 occurrences) document its core European range across the Alps and Carpathians (France, Switzerland, Italy, Austria, Germany, Slovenia, Poland, Romania, Slovakia, Spain, Croatia) extending eastward into Ukraine and disjunctly to the Sino-Himalayan region (China, India). It is a calcicole of stony alpine meadows, rocky ledges and screes, typically between 1,700 and 4,000 m. The Himalayan population photographed at Samiti Lake in Sikkim at 4,000 m in July illustrates the upper elevational reach of the species complex in seasonally snow-fed alpine turf.

The plant forms compact tufts from a woody caudex, with erect unbranched stems 5–20 cm tall arising from basal leaf rosettes. Leaves are lanceolate to oblong, grey-green and densely white-tomentose on both surfaces, the indumentum of interwoven hairs reducing transpiration and reflecting intense UV radiation. The conspicuous "flower" is in fact a pseudanthium: several small yellowish-white capitula are surrounded by a stellate arrangement of densely woolly, snow-white bracts forming the characteristic star. Each capitulum contains tubular florets, with marginal female and central functionally male florets; the achenes are small and bear a pappus of barbellate bristles.

The Leontopodium nivale complex is taxonomically intricate, with subsp. alpinum often treated as the type European edelweiss and frequently confused with Himalayan congeners such as L. himalayanum. A celebrated symbol of alpine flora, it is legally protected in several European countries against collection, and its woolly indumentum is a textbook example of adaptation to cold, high-irradiance environments.

Author: (Cass.) Greuter · Sources: POWO (Kew), Flora of China Vol. 20-21, Flora Europaea Vol. 4, GBIF, Handel-Mazzetti: Monographie der Gattung Leontopodium (1928)

Asteraceae

Ligularia amplexicaulis

by G & P Jürgens

DC.

Garw 2005 33 ergebnis ergebnis
India Garhwal Valley of flowers 3050m, September
Garw 2005 34 ergebnis ergebnis
India Garhwal Valley of flowers 3050m, September

Description

Ligularia amplexicaulis DC. is a robust Himalayan composite ranging from Pakistan and northern India through Nepal and Bhutan into southwestern China (gbif_countries: NP, IN, BT, CN, PK, plus collection records coded US and CA), with around 271 occurrences in GBIF. It is characteristic of moist alpine and subalpine meadows, tall-herb slopes, forest clearings and streamside grassland, typically between about 2800 and 4300 m. The photographed plant from the celebrated Valley of Flowers in Garhwal at 3050 m, flowering in September, exemplifies its late-season presence in the rich monsoon-fed herb meadows for which that valley is famous.

It is a stout perennial herb arising from a short rhizome with fibrous roots. The lower leaves are large, reniform to broadly cordate and long-petiolate, the cauline leaves becoming sessile and conspicuously amplexicaul (stem-clasping), as the epithet records. The inflorescence is an elongate terminal raceme of numerous radiate capitula. Each head bears yellow ray florets and yellow tubular disc florets, subtended by a campanulate involucre of few subequal phyllaries; the achenes are glabrous and cylindric, crowned by a pappus of soft barbellate bristles.

Ligularia (tribe Senecioneae) is a large Sino-Himalayan genus centred on the eastern alpine flora, and the present species is part of the racemose-headed Himalayan assemblage. Like its relatives it accumulates pyrrolizidine alkaloids, deterring herbivory and rendering it of pharmacological and toxicological interest. The species is widespread and locally abundant in protected alpine areas such as Nanda Devi and the Valley of Flowers, and is not considered threatened.

Author: DC. · Sources: POWO (Kew), GBIF, Flora of China (Asteraceae: Senecioneae), Flora of Bhutan, Annotated checklist of the flowering plants of Nepal