Tag:

Summer

Boraginaceae

Maharanga emodi

by G & P Jürgens

DC.

BhutanPG2011 99 scaled
Bhutan Chogcham, 3400m June
Nepal 99 42 ergebnis
West-Nepal Nyor Gaad 2800m, June

Description

Maharanga emodi is a striking bristly herb of the Himalaya and adjacent China, recorded from Nepal, India, China and Bhutan, with GBIF documenting 170 occurrences. It grows on open alpine and subalpine slopes, rocky banks, grazed meadows and cliff ledges. The images derive from Chogcham in Bhutan at about 3400 m and from Nyor Gaad in West Nepal at about 2800 m, both flowering in June, illustrating its mid- to high-montane range across the chain.

The plant is a perennial with a stout rootstock and erect to ascending stems densely clothed in stiff, often pustular-based hairs. The leaves are oblong to oblanceolate, rough-hairy and crowded below. The flowers are pendent in nodding, branched cymes; the corolla is tubular and somewhat inflated or urceolate, contracted at the mouth, typically yellow flushed with purple or wine-red, with the anthers connivent around the exserted style, a floral form distinctive of the genus.

GBIF treats this binomial as a synonym while listing Maharanga emodi as the accepted name, reflecting nomenclatural movement between Maharanga and Onosma. Maharanga is a small Himalayan segregate of Onosma defined largely by its inflated corolla and connate stamens, and its taxonomic independence has been debated; some authors subsume it within Onosma. Like its relatives the roots contain red dye pigments. Its showy flowers and montane habitat make it a conspicuous element of Himalayan slope vegetation.

Author: DC. · Sources: POWO (Kew), GBIF, Flora of Nepal, Flora of Bhutan, Riedl's treatment of Boraginaceae

Asparagaceae

Maianthemum purpureum

by G & P Jürgens

(Wall.) LaFrankie

BhutanPG2011 169
Bhutan Tsochenchen, 3550m June
Nepal 99 100 ergebnis
West-Nepal South of Rara-Lake 3600m, June

Description

Maianthemum purpureum (Wall.) LaFrankie is a montane false Solomon's-seal of the Himalaya and southwestern China, with about 673 records from China, Nepal, India, Bhutan, Myanmar and Taiwan. It grows in shaded, humus-rich situations: fir and rhododendron forest, subalpine scrub, mossy stream banks and the edges of high meadows, typically in the upper temperate to subalpine belt around 3000 to 3800 m. The plants photographed at Tsochenchen in Bhutan at 3550 m and south of Rara Lake in West Nepal at 3600 m, both in June, are typical of its cool, moist, snow-influenced woodland and forest-margin habitats.

The species is an erect rhizomatous perennial arising from a creeping, somewhat fleshy rhizome. The simple, unbranched stem bears several alternate, sessile to subsessile, ovate to elliptic leaves with conspicuous parallel venation. The inflorescence is a terminal raceme or narrow panicle of small flowers; in contrast to typical white-flowered Maianthemum, the perianth here is purple to dull reddish, with six free or shortly united tepals and six stamens. The fruit is a small globose berry that ripens red to purplish-black, each containing one to few seeds.

Long placed in Smilacina (and Tovaria), the species was transferred to a broadly circumscribed Maianthemum, which subsumes those segregate genera on molecular and morphological grounds. Its purple-flowered condition is notable within the predominantly white-flowered genus. The nectariferous flowers attract small insects, and the fleshy berries are dispersed by birds. Widespread across the Sino-Himalayan region and locally common in suitable forest, M. purpureum is not considered threatened, although it depends on the persistence of undisturbed montane woodland.

Author: (Wall.) LaFrankie · Sources: POWO (Kew), GBIF, Flora of China, Flora of Bhutan, Flora of Nepal, LaFrankie's revision of Maianthemum (incl. Smilacina)

Orchideaceae

Malaxis muscifera

by GP Jürgens

(Lindl.) Kuntze

BhutanPG2011 788 01 01
Bhutan Chele-La, 3750m July

Description

Malaxis muscifera is a small terrestrial orchid widespread through the Himalaya and adjacent China, recorded from Nepal, India, China, Bhutan, Pakistan and Myanmar, with some 237 occurrences in GBIF. It grows in moist temperate to subalpine forest and forest margins, among shaded humus, mossy banks and grassy clearings, broadly between about 2,500 and 4,000 m. The plant figured was photographed at Chele-La in western Bhutan at around 3,750 m in July, when the species flowers during the summer monsoon in the cool, damp conditions of the upper conifer and Rhododendron belt.

It is a slender, glabrous perennial herb arising from a small pseudobulb-like, fleshy stem-base. There are usually two, occasionally more, ovate to elliptic, plicate-veined leaves clasping the lower stem by their sheathing bases. The erect stem terminates in a long, dense, many-flowered raceme of minute greenish to yellowish flowers. The narrow sepals and the very slender, thread-like petals spread or reflex, giving the inflorescence its characteristic spidery aspect; the labellum is uppermost through resupination, broadly cordate-auriculate at the base and abruptly contracted to a narrow apex, embracing the short column. The fruit is a small ellipsoid capsule with numerous dust-like seeds.

Malaxis is a large, mainly tropical and warm-temperate genus, and M. muscifera, with its narrowly filiform petals, is among the most distinctive Himalayan representatives, the epithet alluding to the fly-like floral appearance. The flowers are pollinated by small insects, and germination requires mycorrhizal fungi. Frequent and locally abundant across the Himalaya, it is of low conservation concern, though, as an orchid, it is covered by the CITES Appendix II listing of the family.

Author: (Lindl.) Kuntze · Sources: POWO (Kew), GBIF, Flora of China, Flora of Bhutan, Annotated Checklist of the Flowering Plants of Nepal, Pridgeon et al. Genera Orchidacearum

Euphorbiaceae

Mallotus nepalensis

by GP Jürgens

Müll.Arg.

Bh09 231 P387
Bhutan Wangdi, 1650m June
Bh09 380 G601
Bhutan Wangdi, 1650m June
Sikkim 2006 255 ergebnis
India West-Bengal, Lingza 1250m, May

Description

Mallotus nepalensis is a small tree or shrub of the subtropical to lower-temperate Himalaya and its eastern extensions, documented by GBIF (972 occurrences) across China, Nepal, India, Bhutan, Myanmar, Taiwan, Vietnam, Japan and Pakistan. It grows in secondary forest, scrub, forest edges and along streams and clearings, characteristically as a light-demanding component of disturbed and successional vegetation, mostly between about 800 and 2000 m. The collections from Wangdi in Bhutan at 1650 m in June and from Lingza in West Bengal at 1250 m in May illustrate its montane-subtropical distribution along the southern Himalayan front.

The plant is a deciduous shrub or small tree whose young branchlets, petioles and inflorescences are clothed in stellate hairs and reddish, granular peltate glands typical of the genus. The leaves are alternate, long-petiolate, broadly ovate to deltoid, often with two glands at the blade base and a pair of basal lateral nerves, the lower surface pale and gland-dotted. The plant is dioecious; minute apetalous flowers are borne in slender terminal and axillary racemes or panicles, the staminate with numerous stamens, the pistillate with a softly spiny ovary. The fruit is a globose, three-lobed capsule beset with soft processes and covered in red glandular powder.

Mallotus belongs to subfamily Acalyphoideae of the Euphorbiaceae and is characterised by its stellate indumentum and resinous peltate glands, the latter yielding the dye-like powder for which several species are known. Ecologically it behaves as a pioneer, regenerating freely in gaps and along disturbed margins, its capsular fruit and small seeds aiding rapid colonisation. The species is common and of no conservation concern.

Author: Müll.Arg. · Sources: POWO (Kew), GBIF, Flora of China Vol. 11, Flora of Bhutan Vol. 1, Flora of Nepal, Airy Shaw Euphorbiaceae of the Indian subcontinent

Lamiaceae

Marmoritis nivalis 

by G & P Jürgens

(Jacqu. ex Benth.) Hedge

Nepal 99 790 ergebnis
West-Nepal Baga-La-Phedi 4460m, July

Description

Marmoritis nivalis is a high-alpine herb of the Himalaya, recorded from India, Nepal, Bhutan and China; with only about 36 georeferenced records it is a sparsely collected, narrowly montane species. As the epithet "nivalis" indicates, it is a plant of the snow line, growing on open screes, stony alpine turf, moraine and rocky slopes typically between 4000 and 5000 m. The photographed individual from Baga-La-Phedi in West Nepal at 4460 m, flowering in July, exemplifies its occurrence in the cushion-and-scree zone above the timberline, where the short summer growing season coincides with the monsoon.

It is a low, mat-forming or tufted perennial herb arising from a branched woody caudex and slender rhizome, the stems short, ascending and grey-pubescent. The leaves are small, opposite, ovate to orbicular, crenate, thick and densely hairy, an adaptation to cold, exposed, high-radiation habitats. The flowers are crowded in compact terminal heads subtended by leafy bracts; each corolla is bilabiate, pale violet to bluish, with a notched upper lip and a spreading three-lobed lower lip. The calyx is tubular and conspicuously veined, and the four stamens lie under the upper lip. The fruit consists of four small nutlets enclosed in the persistent calyx.

This taxon is treated by GBIF as a synonym, with Marmoritis nivalis itself as the accepted name (combination by Hedge, based on Bentham's earlier name in Dracocephalum/Nepeta). Marmoritis is a small Sino-Himalayan genus segregated from the Nepeta alliance of the Lamiaceae. The cushion habit and dense indumentum are classic high-alpine adaptations. Pollination is by hardy bumblebees active in the brief alpine summer. Owing to its restricted, high-elevation range it warrants monitoring, though no formal threat status is established.

Author: (Jacqu. ex Benth.) Hedge · Sources: POWO (Kew), GBIF, Flora of Nepal, Flora of China, Flora of Bhutan, Hedge's treatment of Himalayan Nepeteae (Lamiaceae)

Papaveraceae

Meconopsis aculeata

by G & P Jürgens

Royle

Garh 07 548 ergebnis
Garh 07 447 ergebnis
India Garhwal Khiar Kutti 3850m, June

Description

Meconopsis aculeata is a Western Himalayan endemic of the high mountains of northern India, Pakistan and adjacent China (Tibet), extending marginally into Nepal; its roughly 249 GBIF records concentrate in the north-western ranges, with cultivated and herbarium material accounting for the scattered occurrences in Great Britain, Canada and Egypt. It grows on open alpine slopes, moist screes, rock crevices and moraine margins, typically between 3000 and 4500 m. The photographed population at Khiar Kutti in the Garhwal Himalaya of India, at 3850 m in June, is characteristic of its early-summer flowering on damp, stony alpine meadow.

It is a monocarpic or short-lived perennial herb forming a stout taproot and a basal rosette. The whole plant is densely armed with stiff, yellowish bristles and prickle-like setae, giving it its epithet aculeata. Leaves are oblong to oblanceolate, pinnately lobed to deeply pinnatifid, narrowed to a winged petiole. The flowering stem bears several large, nodding to spreading flowers; the four petals are clear sky-blue to purplish-blue, broadly obovate and crinkled. Stamens are numerous; the bristly ovary ripens into an ellipsoid, setose capsule dehiscing by short apical valves to release many small seeds.

Within Meconopsis, this species belongs to the blue-flowered, bristly poppies and is among the most widespread of the blue Himalayan poppies west of Nepal. Its dense armature is an adaptation to grazing and harsh exposure, and like many monocarpic Meconopsis it depends on a multi-year rosette to accumulate resources before a single flowering episode. It remains locally frequent and is not formally assessed as threatened, though habitat degradation and collection pressure are localised concerns.

Author: Royle · Sources: POWO (Kew), GBIF, Flora of Pakistan, Flora of British India, The Genus Meconopsis (Grey-Wilson monograph)

Papaveraceae

Meconopsis bella

by G & P Jürgens

Prain

BhutanPG2011 198
Bhutan Tsochenchen, 3650m June
BhutanPG2011 206
Bhutan Tsochenchen, 3650m June

Description

Meconopsis bella is a dwarf, exquisite blue poppy endemic to the eastern Himalaya, recorded from Nepal, northern India, Bhutan and adjacent southern China, the 79 GBIF occurrences reflecting its narrow range and the inaccessibility of its habitat. It grows in cool, moist, sheltered situations at high elevation, characteristically rooted in crevices of rock faces, cliff ledges, moraines and shaded banks between roughly 3300 and 4500 m. The photographed plant was found at Tsochenchen in Bhutan at 3650 m in June, the early-summer flowering season when it blooms amid the monsoon mists of the subalpine zone.

It is a small, slender, tap-rooted perennial lacking a leafy stem, the few leaves forming a basal tuft; the blades are obovate and pinnately lobed or coarsely toothed, on long slender petioles, bearing scattered fine hairs. The flowers are solitary on long, thread-like leafless scapes that arise directly from the rootstock and nod gracefully. Each flower has four broad, satiny petals in pale sky-blue to lavender or, in some forms, soft pink-mauve, surrounding a boss of yellow stamens and a prominent ovary tipped by a clavate stigma. The fruit is a narrow erect capsule dehiscing apically to shed numerous small seeds.

The genus Meconopsis is the celebrated assemblage of Asiatic blue poppies, and M. bella is among the smallest and most delicate, belonging to the crevice-dwelling group. Its scattered, cliff-bound populations and exacting requirements for cool humid air and sharp drainage render it both horticulturally prized and intrinsically vulnerable to collection and climatic shifts. Although not formally Red-Listed, its restricted eastern Himalayan endemism warrants conservation attention, and seed-grown stock underpins its limited cultivation in specialist gardens.

Author: Prain · Sources: POWO (Kew), GBIF, Flora of Bhutan, Grey-Wilson: The Genus Meconopsis, Flora of China

Papaveraceae

Meconopsis chankheliensis

by G & P Jürgens

Grey-Wilson

Nepal 99 248 ergebnis
West-Nepal Tallekh 3155m, June
Nepal 99 265 ergebnis
West-Nepal Tallekh 3155m, June
Nepal 99 266 ergebnis
West-Nepal Tallekh 3155m, June

Description

Meconopsis chankheliensis is a narrowly endemic poppy of the western Himalaya, presently documented only from Nepal, where the comparatively small number of records (18 georeferenced occurrences) reflects both its restricted range and the remoteness of its habitat. It is a plant of moist, sheltered situations in the upper temperate and subalpine zones, typically growing in damp meadows, on humus-rich banks and among rocks beside seepages and streamsides. The photographed material from Tallekh in West Nepal, at about 3155 m and flowering in June, is characteristic of the moderate elevations at which the species comes into bloom following the onset of the early summer monsoon moisture.

The plant is a tufted herb arising from a basal rosette of leaves and a stout taproot. The foliage is more or less covered in spreading bristly hairs; the leaves are oblanceolate to obovate, narrowed to a winged petiole, with entire to lobed or coarsely toothed margins. Flowering stems are erect and bristly, bearing one to several flowers. The blooms are bowl-shaped and showy, with four (occasionally more) broad, satiny petals in shades of blue to purplish-blue, surrounding a dense boss of numerous stamens with slender filaments and a short style crowned by a lobed stigma. The fruit is an elongate, bristly capsule dehiscing by apical valves to release many small seeds.

Meconopsis is a genus of the Sino-Himalayan region whose blue poppies have attracted intense horticultural and taxonomic attention, much of it through the monographic work of Christopher Grey-Wilson, whose name is associated with this taxon. As a localised Nepalese endemic restricted to a single mountain region, M. chankheliensis is of conservation concern, vulnerable to grazing pressure, habitat disturbance and the climatic shifts increasingly affecting Himalayan alpine floras. Its narrow distribution makes accurate field documentation and protection of known populations particularly important.

Author: Grey-Wilson · Sources: POWO (Kew), GBIF, Flora of Nepal, Annotated Checklist of the Flowering Plants of Nepal, Grey-Wilson: The Genus Meconopsis (monograph)

Papaveraceae

Meconopsis grandis

by GP Jürgens

Prain

Nepal 97 187 G369
Nepal Tayla-La 4050m April
Nepal 99 107 ergebnis
West-Nepal South of Rara-Lake 3600m, June
Nepal 99 280 ergebnis
West-Nepal Kalomura 4100m, June
Nepal 99 281 ergebnis
West-Nepal Kalomura 4100m, June

Description

Meconopsis grandis is a high-altitude perennial of the eastern and central Himalaya, distributed across Nepal, northern India, Sikkim, Bhutan and adjacent southern Tibet (China), with cultivated and naturalised records reflected in the European data (United Kingdom, Belgium, Norway, Sweden). GBIF documents some 266 occurrences, a figure inflated by its long horticultural history as one of the celebrated "blue poppies." In its native range it grows in moist alpine meadows, open scrub, the margins of Rhododendron thickets and along streamsides on humus-rich, well-drained acidic soils. The collections illustrated span the snowmelt belt between roughly 3600 and 4100 m, from Tayla-La in central Nepal at 4050 m in April to the slopes south of Rara Lake at 3600 m and Kalomura at 4100 m in June, where it flowers as the monsoon advances.

It forms a stout, deciduous perennial from a thick rootstock, with a basal rosette of oblanceolate to elliptic leaves clothed in rust-coloured to fulvous bristly hairs, the blades narrowing into a winged petiole and often shallowly toothed. The flowering scape is erect, bristly and bears one to several large nodding flowers, each typically with four (sometimes more) broad, satiny, overlapping petals of intense azure to violet-blue, occasionally purplish. Numerous golden stamens surround a stout ovary crowned by a short, well-defined style and capitate to clavate stigma. The fruit is an elongate, ellipsoid to narrowly ovoid capsule, bristly-hairy, dehiscing by short apical valves to release small seeds.

Meconopsis grandis belongs to a taxonomically intricate complex of perennial blue poppies and has historically hybridised in cultivation, giving rise to the widely grown Meconopsis × sheldonii. Variation in flower colour, leaf indumentum and monocarpic versus polycarpic habit has prompted recognition of several subspecies and forms by different authors. Ecologically it is pollinated chiefly by bumblebees attracted to its conspicuous corolla. Though not formally assessed as threatened across its wide range, populations are localised and sensitive to grazing pressure, trampling and the warming of alpine habitats, making it a flagship species for Himalayan montane conservation.

Author: Prain · Sources: POWO (Kew), GBIF, Flora of Bhutan, Flora of Nepal, Annotated Checklist of the Flowering Plants of Nepal, Christopher Grey-Wilson – The Genus Meconopsis

Papaveraceae

Meconopsis horridula

by GP Jürgens

Hook.f. & Thomson

BhutanPG2011 552 1
Bhutan Ludlow-Sherriff-Trail Dem-Tso, 4650m July
BhutanPG2011 664
Bhutan Ludlow-Sherriff-Trail Dem-Tso, 4700m July
Nepal 99 328 ergebnis
West-Nepal Kalmura 4300m, June

Description

Meconopsis horridula is one of the most widespread of the blue poppies, distributed across the high Himalaya and the Tibetan plateau through China, Nepal, Bhutan and India, with a large dataset of around 1472 occurrences (records reported from cultivation in Britain, Canada and elsewhere reflect its horticultural celebrity). It inhabits open stony slopes, scree, moraine, rock crevices and high alpine turf, typically between about 4000 and 5000 m. The photographed plants from Dem-Tso on the Ludlow-Sherriff Trail in Bhutan at 4650–4700 m in July, and from Kalmura in West Nepal at 4300 m in June, are characteristic of its preference for exposed, well-drained high-elevation ground.

It is a monocarpic herb, the whole plant densely armed with stiff, spreading, pale to purplish spines. The leaves form a basal rosette and are oblanceolate, entire to lobed and likewise spiny on both surfaces and along the petiole. Flowering stems bear nodding, saucer-shaped flowers, solitary on long scapes or several on a leafy raceme; the four (to six) petals are sky-blue to purple or occasionally reddish, surrounding a boss of yellow stamens and a stout ovary topped by a short style. The fruit is an erect, narrowly ellipsoid, bristly capsule dehiscing by apical valves to release numerous small seeds.

Meconopsis horridula, described by Hooker and Thomson, has long served as a catch-all for variable spiny blue poppies, and modern revisions have segregated several taxa (such as M. racemosa and M. prattii) formerly lumped under it. Being monocarpic, individuals flower once and die after seed-set. Though locally common, the genus is subject to over-collection of seed and pressure from trekking and grazing across its range.

Author: Hook.f. & Thomson · Sources: POWO (Kew), GBIF, Flora of China (Papaveraceae), Annotated Checklist of the Flowering Plants of Nepal, Grierson & Long Flora of Bhutan, Christopher Grey-Wilson's monograph The Genus Meconopsis